Equations describing model dynamics within the nth growing season.

Turn-over of resource, development of juveniles, allocation of energy storage for reproduction by adults and mortality of consumer individuals are assumed to take place continuously throughout the growing season. Individuals will experience starvation mortality rate when their net biomass productivity is not sufficient to cover their maintenance requirements, which occurs when ν_j(R) < 0 or ν_a(R) < 0. Maturation of juveniles and allocation to energy storage by adults cease when juveniles or adults starve, respectively.