Experimental and BF analysis of CO patterns in tomato.

(A, B) Experimental CoC data for chromosomes with shorter or longer SC lengths (2–4 and 5–11, respectively). Curves for the two groups are offset when inter-interval distance is expressed in units of Mb (genomic distance) (Panel A) and are overlapping when distance is expressed in units of µm SC (physical distance) (Panel B). (C, D) BF best-fit simulations for both groups of chromosomes using the physical distance metric (µm SC length) and adjusting precursor number as required to give ∼20 precursors per CO for each type of chromosome. This relationship is based on observation of ∼280 Rad51 foci [69] and 15 Mlh1 foci genome wide [38]. Interestingly, this relationship further implies that N is proportional to SC length.