[Meredith et al. (2011) molecular tree used for ancestral state reconstruction - node numbers correspond to predicted anceestors (PAs) in other trees]
((((((Orycteropus,(Rhynchocyon,(Amblysomus,Echinops)'45')'44')'43',(Trichechus,(Loxodonta,Procavia)'42')'41')'40',(Dasypus,Tamandua)'39')'38',(((((Equus,(Lama,(Sus,(Bos,(Hippopotamus,(Caperea,Tursiops)'15')'14')'13')'12')'11')'10',((Myotis,(Pteronotus,(Nycteris,Saccopteryx)'19')'18')'17',(Pteropus,Rhinopoma)'20')'16')'9',(Manis,(Canis,Felis)'22')'21')'8',(Solenodon,(Erinaceus,(Sorex,Talpa)'25')'24')'23')'7',(((Oryctolagus,((Castor,Rattus)'30',(Cavia,Ictidomys)'31')'29')'28',(Ptilocercus,Tupaia)'32')'27',((Cynocephalus,Galeopterus)'34',(Lemur,(Tarsius,(Homo,Saimiri)'37')'36')'35')'33')'26')'6')'5',(Didelphis,Dromiciops)'4')'3',(Ornithorhynchus,Tachyglossus)'2')'1';

[Maximum parsimony analyses]

[no PAs: nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,(((Eomaia_scansoria:0.000000,(((((Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.522000:0.000000)0.830000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.848000:0.000000)0.232000:0.000000)0.442000:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.468000:0.000000)0.190000:0.000000,(((Protungulatum_donnae:0.000000,((Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.292000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.348000:0.000000)0.092000:0.000000)0.274000:0.000000,((Thomashuxleya_externa:0.000000,((Procavia_capensis:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.992000:0.000000)0.080000:0.000000)0.542000:0.000000)0.072000:0.000000)0.010000:0.000000)0.024000:0.000000)0.026000:0.000000)0.012000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.972000:0.000000)0.010000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.590000:0.000000)0.250000:0.000000)0.006000:0.000000,(Carodnia_vieirai:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.116000:0.000000)0.380000:0.000000)0.518000:0.000000)0.012000:0.000000)0.180000:0.000000,((((Rattus_norvegicus:0.000000,(Ictidomys_tridecemlineatus:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.874000:0.000000)0.274000:0.000000)0.154000:0.000000,(Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.132000:0.000000)0.246000:0.000000,((Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.888000:0.000000)1.000000:0.000000,(Rhombomylus_turpanensis:0.000000,Tribosphenomys_minutus:0.000000)0.014000:0.000000)0.294000:0.000000)0.996000:0.000000,(Orycteropus_afer:0.000000,((Dasypus_novemcinctus:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.996000:0.000000)0.738000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.888000:0.000000)0.510000:0.000000)0.622000:0.000000)0.022000:0.000000)0.002000:0.000000)0.004000:0.000000)0.006000:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)0.998000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.814000:0.000000)1.000000:0.000000)0.008000:0.000000,((((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.434000:0.000000,(Notharctus_tenebrosus:0.000000,(Lemur_catta:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.980000:0.000000)0.570000:0.000000)0.242000:0.000000)1.000000:0.000000)0.534000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)0.996000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.688000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.762000:0.000000)0.970000:0.000000)1.000000:0.000000)0.988000:0.000000)1.000000:0.000000)0.208000:0.000000)0.066000:0.000000)0.604000:0.000000,(Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.902000:0.000000)0.296000:0.000000)0.144000:0.000000,(Dromiciops_gliroides:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.896000:0.000000)1.000000:0.000000)0.394000:0.000000)1.000000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.858000:0.000000)0.970000:0.000000,(Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)1.000000:0.000000)0.998000:0.000000)1.000000:0.000000)2.000000:0.000000);

[no PAs: nodal values = "transfer bootstrap expectation" values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,(((Eomaia_scansoria:0.000000,(((((Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.522000:0.000000)0.879000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.848000:0.000000)0.658000:0.000000)0.693600:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.468000:0.000000)0.574286:0.000000,(((Protungulatum_donnae:0.000000,((Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.292000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.348000:0.000000)0.370667:0.000000)0.433000:0.000000,((Thomashuxleya_externa:0.000000,((Procavia_capensis:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.992000:0.000000)0.538000:0.000000)0.726667:0.000000)0.568500:0.000000)0.461200:0.000000)0.397667:0.000000)0.358000:0.000000)0.341750:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.972000:0.000000)0.386200:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.590000:0.000000)0.467833:0.000000)0.454308:0.000000,(Carodnia_vieirai:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.116000:0.000000)0.552000:0.000000)0.642667:0.000000)0.448000:0.000000)0.447222:0.000000,((((Rattus_norvegicus:0.000000,(Ictidomys_tridecemlineatus:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.874000:0.000000)0.608000:0.000000)0.502667:0.000000,(Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.132000:0.000000)0.529600:0.000000,((Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.888000:0.000000)1.000000:0.000000,(Rhombomylus_turpanensis:0.000000,Tribosphenomys_minutus:0.000000)0.014000:0.000000)0.725500:0.000000)0.997800:0.000000,(Orycteropus_afer:0.000000,((Dasypus_novemcinctus:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.996000:0.000000)0.869000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.888000:0.000000)0.823000:0.000000)0.848400:0.000000)0.633375:0.000000)0.385600:0.000000)0.381150:0.000000)0.390488:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)0.998000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.938000:0.000000)1.000000:0.000000)0.395789:0.000000,((((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.622667:0.000000,(Notharctus_tenebrosus:0.000000,(Lemur_catta:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.980000:0.000000)0.775000:0.000000)0.747333:0.000000)1.000000:0.000000)0.776500:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)0.996000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.688000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.762000:0.000000)0.978000:0.000000)1.000000:0.000000)0.998000:0.000000)1.000000:0.000000)0.585250:0.000000)0.545524:0.000000)0.816308:0.000000,(Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.902000:0.000000)0.619000:0.000000)0.720400:0.000000,(Dromiciops_gliroides:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.896000:0.000000)1.000000:0.000000)0.913429:0.000000)1.000000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.858000:0.000000)0.985000:0.000000,(Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)1.000000:0.000000)0.999000:0.000000)1.000000:0.000000)-1.#IND00:0.000000);

[“all characters” PAs corrected for the O'Leary et al. character ontology: 24 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA2,(Tachyglossus_aculeatus,PA1))),(((PA3,((PA5,((PA40,((PA43,(Orycteropus_afer,((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))))),((PA6,((PA7,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(((PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15)))))),(Sus_scrofa,PA12)),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(((Erinaceus_europaeus,PA24),(Talpa_europaea,(Sorex_araneus,PA25))),(Solenodon_paradoxus,PA23)))),((PA28,((PA29,(((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(PA33,((PA35,(Tarsius_syrichta,(PA36,(PA37,(Homo_sapiens,Saimiri_sciureus))))),(Lemur_catta,Notharctus_tenebrosus)))),(PA26,(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA2,(Tachyglossus_aculeatus,PA1))),(((PA3,((PA5,((PA40,((PA43,(Orycteropus_afer,((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))))),((PA6,((PA7,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12)),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA23,(Solenodon_paradoxus,((Erinaceus_europaeus,PA24),(Sorex_araneus,(Talpa_europaea,PA25))))))),((PA28,((PA29,((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(PA33,((PA35,((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36))),(Lemur_catta,Notharctus_tenebrosus)))),(PA26,(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA2,(Tachyglossus_aculeatus,PA1))),(((PA3,((PA5,((PA40,((PA43,(Orycteropus_afer,((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla)))))),((PA6,((PA7,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12)),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA23,(Solenodon_paradoxus,((Erinaceus_europaeus,PA24),(Sorex_araneus,(Talpa_europaea,PA25))))))),((PA28,((PA29,(((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA27,(PA26,((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),(PA33,((PA35,((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36))),(Lemur_catta,Notharctus_tenebrosus)))))),(Ptilocercus_lowii,(Tupaia_glis,PA32))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),((((((PA40,((((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))),(Orycteropus_afer,PA43)),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA5,(((PA7,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,((PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus)))))),(Sus_scrofa,PA12))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA23,(Solenodon_paradoxus,(Erinaceus_europaeus,(PA24,(Talpa_europaea,(Sorex_araneus,PA25))))))),(PA6,((PA28,((PA29,((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA26,((PA33,((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),(((Homo_sapiens,(Saimiri_sciureus,PA37)),(Tarsius_syrichta,PA36)),(PA35,(Lemur_catta,Notharctus_tenebrosus))))),(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))),(PA3,PA1)),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,(PA5,(((PA38,((PA40,(((PA44,(Amblysomus_hottentotus,(Echinops_telfairi,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(Orycteropus_afer,PA43))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla)))),((PA6,((PA7,((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA12,(Sus_scrofa,(PA13,(Hippopotamus_amphibius,(PA14,((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15)))))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),((Erinaceus_europaeus,(PA24,(Talpa_europaea,(Sorex_araneus,PA25)))),(Solenodon_paradoxus,PA23)))),((PA28,(((PA29,(PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,(PA33,((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),((PA35,(PA36,(Tarsius_syrichta,(Saimiri_sciureus,(Homo_sapiens,PA37))))),(Lemur_catta,Notharctus_tenebrosus))))),(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,(((PA40,((PA43,(Orycteropus_afer,((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA5,((PA6,(PA7,(((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(PA11,(((PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15)))))),(Sus_scrofa,PA12)),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),(Solenodon_paradoxus,(PA23,(Erinaceus_europaeus,(PA24,(PA25,(Sorex_araneus,Talpa_europaea))))))))),((PA28,((PA29,((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(PA33,((PA36,(Tarsius_syrichta,(Homo_sapiens,(Saimiri_sciureus,PA37)))),(PA35,(Lemur_catta,Notharctus_tenebrosus)))))),(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,(PA2,PA1))),(((PA3,(PA5,((PA38,(((PA40,(PA43,(Orycteropus_afer,(PA44,((PA45,(Amblysomus_hottentotus,Echinops_telfairi)),(Leptictis_dakotensis,Rhynchocyon_cirnei)))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),((((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,(Sus_scrofa,(PA12,(PA13,(Hippopotamus_amphibius,(PA14,((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus))))))))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA7,(((Erinaceus_europaeus,PA24),(PA25,(Sorex_araneus,Talpa_europaea))),(Solenodon_paradoxus,PA23)))),(PA6,((PA28,((((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,((PA33,(PA34,(Cynocephalus_volans,Galeopterus_variegatus))),((PA35,(Tarsius_syrichta,(PA36,(Saimiri_sciureus,(Homo_sapiens,PA37))))),(Lemur_catta,Notharctus_tenebrosus)))),(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,(PA2,PA1))),(((PA3,(((PA40,((PA43,(Orycteropus_afer,(PA44,((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,(PA39,(Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA5,((PA6,(PA7,(((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),(((Erinaceus_europaeus,PA24),(Talpa_europaea,(Sorex_araneus,PA25))),(Solenodon_paradoxus,PA23))))),((PA28,((PA29,((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),(PA33,((PA35,(Tarsius_syrichta,(PA36,(Homo_sapiens,(Saimiri_sciureus,PA37))))),(Lemur_catta,Notharctus_tenebrosus))))),(PA27,(Tupaia_glis,(Ptilocercus_lowii,PA32))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((((PA5,(PA38,((PA40,((((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))),(Orycteropus_afer,PA43)),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA39,(Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)))))),((((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,(Sus_scrofa,(PA12,(PA13,(Hippopotamus_amphibius,(PA14,((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus))))))))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA7,(Solenodon_paradoxus,(PA23,((Erinaceus_europaeus,PA24),(Sorex_araneus,(Talpa_europaea,PA25))))))),(PA6,((PA28,((((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA27,((PA26,((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(PA33,((PA35,((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36))),(Lemur_catta,Notharctus_tenebrosus))))),(Tupaia_glis,(Ptilocercus_lowii,PA32)))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((((PA5,(PA38,((PA40,((((PA45,(Amblysomus_hottentotus,Echinops_telfairi)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))),(Orycteropus_afer,PA43)),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))))),((((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,(Sus_scrofa,(PA12,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus))),(Hippopotamus_amphibius,PA14)))))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA7,(PA23,(Solenodon_paradoxus,(Erinaceus_europaeus,(PA24,(PA25,(Sorex_araneus,Talpa_europaea)))))))),(PA6,((PA28,((((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA27,((PA26,((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(PA33,((PA35,(Tarsius_syrichta,(PA36,(Saimiri_sciureus,(Homo_sapiens,PA37))))),(Lemur_catta,Notharctus_tenebrosus))))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA3,(PA1,(PA5,(((PA38,((Orycteropus_afer,(PA43,((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))),(PA6,((PA7,(((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,(PA12,(Sus_scrofa,(PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus))))))))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),(((Erinaceus_europaeus,PA24),(Talpa_europaea,(Sorex_araneus,PA25))),(Solenodon_paradoxus,PA23)))),((PA28,(((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA26,(((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),(PA33,(((Homo_sapiens,(Saimiri_sciureus,PA37)),(Tarsius_syrichta,PA36)),(PA35,(Lemur_catta,Notharctus_tenebrosus))))),(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA3,(PA1,(PA5,((PA38,(((((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))),(Orycteropus_afer,PA43)),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA6,(((PA7,(PA8,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(PA11,((Sus_scrofa,(PA12,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus))),(Hippopotamus_amphibius,PA14))))),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA23,(Solenodon_paradoxus,((Erinaceus_europaeus,PA24),(Sorex_araneus,(Talpa_europaea,PA25)))))),((PA28,((((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,(PA33,((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),(((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36)),(PA35,(Lemur_catta,Notharctus_tenebrosus)))))),(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA3,(PA1,(PA5,(((PA38,((((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))),(Orycteropus_afer,PA43)),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))),(PA6,((PA7,(((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,(Sus_scrofa,(PA12,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15))),(Hippopotamus_amphibius,PA14)))))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),(PA23,(Solenodon_paradoxus,((Erinaceus_europaeus,PA24),(PA25,(Sorex_araneus,Talpa_europaea))))))),((PA28,(((PA29,(PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA26,((PA33,((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),((PA35,(Tarsius_syrichta,(PA36,(Saimiri_sciureus,(Homo_sapiens,PA37))))),(Lemur_catta,Notharctus_tenebrosus)))),(PA27,(Tupaia_glis,(Ptilocercus_lowii,PA32)))))))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((((PA38,(((PA40,(Orycteropus_afer,(PA43,(PA44,((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(Leptictis_dakotensis,Rhynchocyon_cirnei)))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA5,((PA6,(PA7,(((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),(((Erinaceus_europaeus,PA24),(PA25,(Sorex_araneus,Talpa_europaea))),(Solenodon_paradoxus,PA23))))),((PA28,(((PA29,(PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,((PA33,(Cynocephalus_volans,(Galeopterus_variegatus,PA34))),((PA35,(Tarsius_syrichta,(PA36,(Homo_sapiens,(Saimiri_sciureus,PA37))))),(Lemur_catta,Notharctus_tenebrosus)))),(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,(PA5,(((PA38,((Orycteropus_afer,(PA43,(PA44,((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),(PA39,(Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA6,((((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),(PA7,(Solenodon_paradoxus,(PA23,((Erinaceus_europaeus,PA24),(Sorex_araneus,(Talpa_europaea,PA25))))))),((PA28,((((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA27,((PA26,((PA33,(Galeopterus_variegatus,(Cynocephalus_volans,PA34))),((PA35,(PA36,(Tarsius_syrichta,(Saimiri_sciureus,(Homo_sapiens,PA37))))),(Lemur_catta,Notharctus_tenebrosus)))),(Tupaia_glis,(Ptilocercus_lowii,PA32)))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,((PA5,((PA40,((Orycteropus_afer,(PA43,((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla)))))),(((PA7,(PA8,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12)),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(Solenodon_paradoxus,(PA23,(Erinaceus_europaeus,(PA24,(Talpa_europaea,(Sorex_araneus,PA25))))))),(PA6,((PA28,((PA29,((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,(PA33,((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),((PA35,((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36))),(Lemur_catta,Notharctus_tenebrosus))))),(PA27,(PA32,(Ptilocercus_lowii,Tupaia_glis)))))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,((PA5,((PA40,((((PA44,(PA45,(Amblysomus_hottentotus,Echinops_telfairi))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(Orycteropus_afer,PA43)),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA6,((PA7,(((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA12,(Sus_scrofa,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15))),(Hippopotamus_amphibius,PA14))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(Solenodon_paradoxus,(PA23,((Erinaceus_europaeus,PA24),(PA25,(Sorex_araneus,Talpa_europaea))))))),((PA28,(((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA27,(PA26,((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(PA33,((PA35,(Tarsius_syrichta,(PA36,(Homo_sapiens,(Saimiri_sciureus,PA37))))),(Lemur_catta,Notharctus_tenebrosus)))))),(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,((PA5,((PA40,((Orycteropus_afer,(PA43,((PA45,(Amblysomus_hottentotus,Echinops_telfairi)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla)))))),(((PA7,(PA8,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12)),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(Solenodon_paradoxus,(PA23,(Erinaceus_europaeus,(PA24,(PA25,(Sorex_araneus,Talpa_europaea))))))),(PA6,((PA28,(((PA29,(PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,(PA33,((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),((PA35,(Tarsius_syrichta,(PA36,(Homo_sapiens,(Saimiri_sciureus,PA37))))),(Lemur_catta,Notharctus_tenebrosus))))),(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA3,(PA1,((((((PA44,(PA45,(Amblysomus_hottentotus,Echinops_telfairi))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(Orycteropus_afer,PA43)),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA5,(((PA7,((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(PA11,(((PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus)))))),(Sus_scrofa,PA12)),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA23,(Solenodon_paradoxus,(Erinaceus_europaeus,(PA24,(Talpa_europaea,(Sorex_araneus,PA25))))))),(PA6,((PA28,((PA29,(((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),(PA33,((PA36,(Tarsius_syrichta,(Saimiri_sciureus,(Homo_sapiens,PA37)))),(PA35,(Lemur_catta,Notharctus_tenebrosus))))),(PA26,(PA27,(Tupaia_glis,(Ptilocercus_lowii,PA32)))))))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((((PA38,((PA40,((Orycteropus_afer,(PA43,((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA5,(PA6,(((PA7,((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(PA11,((PA12,(Sus_scrofa,(PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(Solenodon_paradoxus,(PA23,(Erinaceus_europaeus,(PA24,(Talpa_europaea,(Sorex_araneus,PA25))))))),((PA28,(((PA29,((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA27,(PA26,((PA33,(Cynocephalus_volans,(Galeopterus_variegatus,PA34))),(PA35,(((Homo_sapiens,(Saimiri_sciureus,PA37)),(Tarsius_syrichta,PA36)),(Lemur_catta,Notharctus_tenebrosus)))))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,((PA5,((PA38,(PA40,((PA43,(Orycteropus_afer,((PA45,(Amblysomus_hottentotus,Echinops_telfairi)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA6,(((PA7,((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,(PA12,(Sus_scrofa,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus))),(Hippopotamus_amphibius,PA14)))))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),((Erinaceus_europaeus,(PA24,(Sorex_araneus,(Talpa_europaea,PA25)))),(Solenodon_paradoxus,PA23))),((PA28,(((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA26,(((PA33,(Galeopterus_variegatus,(Cynocephalus_volans,PA34))),((PA35,((PA37,(Homo_sapiens,Saimiri_sciureus)),(Tarsius_syrichta,PA36))),(Lemur_catta,Notharctus_tenebrosus))),(PA27,(PA32,(Ptilocercus_lowii,Tupaia_glis)))))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,((PA5,((PA38,(PA40,((PA43,(Orycteropus_afer,((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA6,((PA7,(((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),((Erinaceus_europaeus,(PA24,(Sorex_araneus,(Talpa_europaea,PA25)))),(Solenodon_paradoxus,PA23)))),((PA28,(((PA29,(Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus)))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(((PA33,(Galeopterus_variegatus,(Cynocephalus_volans,PA34))),(((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36)),(PA35,(Lemur_catta,Notharctus_tenebrosus)))),(PA26,(PA27,(Tupaia_glis,(Ptilocercus_lowii,PA32)))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,(PA5,((PA38,(((Orycteropus_afer,(PA43,((PA44,(Echinops_telfairi,(Amblysomus_hottentotus,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)))),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA39,(Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla))))),((PA6,(PA7,((PA8,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,(PA12,(Sus_scrofa,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14)))))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),((Erinaceus_europaeus,(PA24,(Sorex_araneus,(Talpa_europaea,PA25)))),(Solenodon_paradoxus,PA23))))),((PA28,(((PA29,(Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus)))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA27,((PA26,(PA33,((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),((PA35,(PA36,(Tarsius_syrichta,(Homo_sapiens,(Saimiri_sciureus,PA37))))),(Lemur_catta,Notharctus_tenebrosus))))),(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,((PA5,(((PA43,(Orycteropus_afer,((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))))),((PA6,((PA8,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(PA11,((PA12,(Sus_scrofa,(PA13,(Hippopotamus_amphibius,(PA14,((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15)))))))),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA7,(Solenodon_paradoxus,(PA23,(Erinaceus_europaeus,(PA24,(Talpa_europaea,(Sorex_araneus,PA25))))))))),((PA28,((PA29,((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA27,((PA26,((PA33,(Cynocephalus_volans,(Galeopterus_variegatus,PA34))),(((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36)),(PA35,(Lemur_catta,Notharctus_tenebrosus))))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));

[“all characters” PAs corrected for the O'Leary et al. character ontology: strict consensus of 24 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000,PA3:0.000000,PA2:0.000000,PA1:0.000000,(PA5:0.000000,(PA38:0.000000,(PA40:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(PA44:0.000000,(Amblysomus_hottentotus:0.000000,Echinops_telfairi:0.000000,PA45:0.000000)0.464000:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.036000:0.000000)0.020000:0.000000)0.020000:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.264000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.128000:0.000000)0.068000:0.000000)0.052000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.376000:0.000000)0.020000:0.000000)0.088000:0.000000)0.000000:0.000000,(Dasypus_novemcinctus:0.000000,PA39:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.992000:0.000000)0.624000:0.000000)0.000000:0.000000,(PA6:0.000000,(PA7:0.000000,(PA8:0.000000,(((((Apheliscus_insidiosus:0.000000,Protungulatum_donnae:0.000000)0.028000:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.324000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.260000:0.000000)0.056000:0.000000)0.132000:0.000000,(Rodhocetus_balochistanensis:0.000000,(Archaeotherium_mortoni:0.000000,Mesonyx_obtusidens:0.000000)0.056000:0.000000)0.096000:0.000000)0.004000:0.000000,((PA10:0.000000,(PA11:0.000000,(Sus_scrofa:0.000000,PA12:0.000000,(PA13:0.000000,(Hippopotamus_amphibius:0.000000,PA14:0.000000,((Artiocetus_clavis:0.000000,Basilosaurus_cetoides:0.000000)0.244000:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000,PA15:0.000000)0.992000:0.000000)0.184000:0.000000)0.176000:0.000000)0.008000:0.000000)0.008000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.020000:0.000000)0.032000:0.000000)0.008000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.424000:0.000000)0.024000:0.000000)0.004000:0.000000,(PA9:0.000000,(((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,(((Myotis_lucifugus:0.000000,PA17:0.000000)0.188000:0.000000,(Pteronotus_parnellii:0.000000,PA18:0.000000)0.120000:0.000000)0.360000:0.000000,((Nycteris_thebaica:0.000000,PA19:0.000000)0.084000:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.392000:0.000000)0.648000:0.000000)0.592000:0.000000)0.416000:0.000000,(PA16:0.000000,(Pteropus_giganteus:0.000000,PA20:0.000000)0.416000:0.000000)0.096000:0.000000)1.000000:0.000000)0.040000:0.000000)0.000000:0.000000,(PA21:0.000000,(((Canis_lupus:0.000000,Felis_silvestris:0.000000,PA22:0.000000)0.788000:0.000000,(Sinopa_rapax:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.956000:0.000000)0.952000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.960000:0.000000)0.060000:0.000000)0.176000:0.000000)0.000000:0.000000,(Solenodon_paradoxus:0.000000,PA23:0.000000,(Erinaceus_europaeus:0.000000,PA24:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000,PA25:0.000000)0.812000:0.000000)0.204000:0.000000)0.156000:0.000000)0.000000:0.000000,((PA28:0.000000,((PA29:0.000000,(Ictidomys_tridecemlineatus:0.000000,PA31:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.164000:0.000000)0.120000:0.000000,(Rattus_norvegicus:0.000000,PA30:0.000000)0.180000:0.000000)0.268000:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.652000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.776000:0.000000)0.996000:0.000000)0.424000:0.000000)0.388000:0.000000)0.364000:0.000000)0.628000:0.000000)1.000000:0.000000,(PA27:0.000000,PA26:0.000000,(PA33:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000,PA34:0.000000)1.000000:0.000000,(PA35:0.000000,(Tarsius_syrichta:0.000000,PA36:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000,PA37:0.000000)1.000000:0.000000)0.572000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000)0.416000:0.000000)1.000000:0.000000)0.844000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000,PA32:0.000000)1.000000:0.000000)0.528000:0.000000)0.840000:0.000000)0.000000:0.000000)0.120000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.976000:0.000000)0.972000:0.000000,((Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000)0.492000:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.312000:0.000000)0.604000:0.000000)0.604000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.984000:0.000000)1.000000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“all characters” PAs corrected for the O'Leary et al. character ontology: strict consensus of 24 MPTs - nodal values = "transfer bootstrap expectation" values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000,PA3:0.000000,PA2:0.000000,PA1:0.000000,(PA5:0.000000,(PA38:0.000000,(PA40:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(PA44:0.000000,(Amblysomus_hottentotus:0.000000,Echinops_telfairi:0.000000,PA45:0.000000)0.510000:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.036000:0.000000)0.268000:0.000000)0.212000:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.264000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.128000:0.000000)0.194000:0.000000)0.258000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.376000:0.000000)0.295333:0.000000)0.326286:0.000000)0.203000:0.000000,(Dasypus_novemcinctus:0.000000,PA39:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.992000:0.000000)0.748000:0.000000)0.193333:0.000000,(PA6:0.000000,(PA7:0.000000,(PA8:0.000000,(((((Apheliscus_insidiosus:0.000000,Protungulatum_donnae:0.000000)0.028000:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.324000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.260000:0.000000)0.252000:0.000000)0.297600:0.000000,(Rodhocetus_balochistanensis:0.000000,(Archaeotherium_mortoni:0.000000,Mesonyx_obtusidens:0.000000)0.056000:0.000000)0.150000:0.000000)0.209000:0.000000,((PA10:0.000000,(PA11:0.000000,(Sus_scrofa:0.000000,PA12:0.000000,(PA13:0.000000,(Hippopotamus_amphibius:0.000000,PA14:0.000000,((Artiocetus_clavis:0.000000,Basilosaurus_cetoides:0.000000)0.244000:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000,PA15:0.000000)0.994000:0.000000)0.627000:0.000000)0.506667:0.000000)0.436000:0.000000)0.346222:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.020000:0.000000)0.369000:0.000000)0.382769:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.424000:0.000000)0.418400:0.000000)0.355667:0.000000,(PA9:0.000000,(((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,(((Myotis_lucifugus:0.000000,PA17:0.000000)0.188000:0.000000,(Pteronotus_parnellii:0.000000,PA18:0.000000)0.120000:0.000000)0.366667:0.000000,((Nycteris_thebaica:0.000000,PA19:0.000000)0.084000:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.392000:0.000000)0.696000:0.000000)0.753143:0.000000)0.832000:0.000000,(PA16:0.000000,(Pteropus_giganteus:0.000000,PA20:0.000000)0.416000:0.000000)0.256000:0.000000)1.000000:0.000000)0.926154:0.000000)0.391895:0.000000,(PA21:0.000000,(((Canis_lupus:0.000000,Felis_silvestris:0.000000,PA22:0.000000)0.788000:0.000000,(Sinopa_rapax:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.978000:0.000000)0.970400:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.960000:0.000000)0.726857:0.000000)0.694500:0.000000)0.319417:0.000000,(Solenodon_paradoxus:0.000000,PA23:0.000000,(Erinaceus_europaeus:0.000000,PA24:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000,PA25:0.000000)0.826000:0.000000)0.567000:0.000000)0.456000:0.000000)0.294571:0.000000,((PA28:0.000000,((PA29:0.000000,(Ictidomys_tridecemlineatus:0.000000,PA31:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.164000:0.000000)0.149333:0.000000,(Rattus_norvegicus:0.000000,PA30:0.000000)0.180000:0.000000)0.324000:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.652000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.776000:0.000000)0.996000:0.000000)0.808000:0.000000)0.762000:0.000000)0.676667:0.000000)0.971385:0.000000)1.000000:0.000000,(PA27:0.000000,PA26:0.000000,(PA33:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000,PA34:0.000000)1.000000:0.000000,(PA35:0.000000,(Tarsius_syrichta:0.000000,PA36:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000,PA37:0.000000)1.000000:0.000000)0.850000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000)0.416000:0.000000)1.000000:0.000000)0.943636:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000,PA32:0.000000)1.000000:0.000000)0.815250:0.000000)0.917161:0.000000)0.193300:0.000000)0.406353:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.976000:0.000000)0.973333:0.000000,((Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000)0.492000:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.312000:0.000000)0.697333:0.000000)0.804000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.984000:0.000000)1.000000:0.000000)1.000000:0.000000));

[“all characters” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: 10 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((PA1,(PA3,((PA5,((PA6,((PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus)))),((PA21,(((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))),(Manis_pentadactyla,Metacheiromys_marshi))),(PA8,((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))))))),((PA27,(PA28,((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,(Cocomys_lingchaensis,Paramys_delicatus)))),((Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)),(Rhombomylus_turpanensis,Tribosphenomys_minutus))))),(PA26,((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))))),((PA38,(PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(Orycteropus_afer,(PA43,(Rhynchocyon_cirnei,PA44)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),(((((PA5,((PA6,((PA7,((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(PA8,((PA21,(((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))),(Manis_pentadactyla,Metacheiromys_marshi))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))))))),((PA28,((Rattus_norvegicus,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(PA33,((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus)))),(PA27,(Ptilocercus_lowii,Tupaia_glis))))))),((PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(Orycteropus_afer,(PA43,(Rhynchocyon_cirnei,PA44))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((PA6,(((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(PA7,((PA8,(PA21,(((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))),(Manis_pentadactyla,Metacheiromys_marshi)))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))),((PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(PA26,(((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus))),(PA27,(Ptilocercus_lowii,Tupaia_glis))))))),(PA5,((PA38,(PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(Orycteropus_afer,(PA43,(Rhynchocyon_cirnei,PA44)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA3,PA1)),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((PA1,(PA3,((PA6,(((PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus)))),((PA21,(((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))),(Manis_pentadactyla,Metacheiromys_marshi))),(PA8,((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))),((PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(PA26,(((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus))),(PA27,(Ptilocercus_lowii,Tupaia_glis))))))),(PA5,((PA38,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA40,(Orycteropus_afer,(PA43,(Rhynchocyon_cirnei,PA44)))))),(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla)))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),(((((((PA6,(PA7,((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),((PA21,(((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))),(Manis_pentadactyla,Metacheiromys_marshi))),(PA8,((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))))),((PA28,((Rattus_norvegicus,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA26,((PA33,((Cynocephalus_volans,Galeopterus_variegatus),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus)))),(PA27,(Ptilocercus_lowii,Tupaia_glis)))))),(PA5,((PA38,(PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA43,(Orycteropus_afer,(Rhynchocyon_cirnei,PA44)))))),(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla)))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((PA1,(PA3,((((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(PA7,((PA8,(PA21,(((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))),(Manis_pentadactyla,Metacheiromys_marshi)))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))),(PA6,((PA26,(PA27,(PA28,((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,(Cocomys_lingchaensis,Paramys_delicatus)))),((Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)),(Rhombomylus_turpanensis,Tribosphenomys_minutus)))))),((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))),(PA5,((PA38,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA40,((Orycteropus_afer,PA43),(Rhynchocyon_cirnei,PA44))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((PA1,(PA3,((PA5,((PA6,((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(PA7,((PA8,(PA21,(((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))),(Manis_pentadactyla,Metacheiromys_marshi)))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))))))),((PA26,(PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))))),((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))),(((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA40,(Orycteropus_afer,(PA43,(Rhynchocyon_cirnei,PA44))))),(PA38,(PA39,(Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((PA6,(((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(PA7,((PA21,(((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))),(Manis_pentadactyla,Metacheiromys_marshi))),(PA8,((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))))))),((PA27,(PA28,((Rattus_norvegicus,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(PA26,((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))))),(PA5,((PA38,(PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA43,(Orycteropus_afer,(Rhynchocyon_cirnei,PA44)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA3,PA1)),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((PA1,(PA3,((PA5,((PA6,((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(PA7,(PA8,((PA21,(((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))),(Manis_pentadactyla,Metacheiromys_marshi))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))))),(PA26,((PA27,(PA28,((Rattus_norvegicus,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))))),(((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA40,((Orycteropus_afer,PA43),(Rhynchocyon_cirnei,PA44)))),(PA38,(PA39,(Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),(((((PA5,(((PA7,((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),((PA8,(PA21,(((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))),(Manis_pentadactyla,Metacheiromys_marshi)))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))),(PA6,((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA26,((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))))),(((PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(PA43,(Orycteropus_afer,(Rhynchocyon_cirnei,PA44)))),(PA38,(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));

[“all characters” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: strict consensus of 10 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.896000:0.000000,((((PA3:0.000000,PA1:0.000000,(PA5:0.000000,(PA6:0.000000,(PA7:0.000000,(Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.440000:0.000000)0.792000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.744000:0.000000)0.236000:0.000000)0.296000:0.000000,(PA8:0.000000,(PA21:0.000000,(((Leptictis_dakotensis:0.000000,(Protungulatum_donnae:0.000000,(Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.368000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.272000:0.000000)0.096000:0.000000)0.132000:0.000000)0.228000:0.000000)0.032000:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)0.792000:0.000000,(Sinopa_rapax:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.340000:0.000000)1.000000:0.000000)0.004000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.908000:0.000000)0.000000:0.000000)0.000000:0.000000,((PA9:0.000000,(PA10:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.988000:0.000000)0.224000:0.000000)0.784000:0.000000)0.328000:0.000000)0.220000:0.000000)0.256000:0.000000)0.336000:0.000000)0.268000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.928000:0.000000)0.244000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.616000:0.000000)0.140000:0.000000)0.592000:0.000000)0.064000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.688000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.776000:0.000000)0.984000:0.000000)1.000000:0.000000)0.992000:0.000000)1.000000:0.000000)0.160000:0.000000)0.000000:0.000000)0.000000:0.000000,(PA27:0.000000,PA33:0.000000,PA26:0.000000,(PA28:0.000000,(Ictidomys_tridecemlineatus:0.000000,Rattus_norvegicus:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.904000:0.000000,(Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.364000:0.000000,(Rhombomylus_turpanensis:0.000000,Tribosphenomys_minutus:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.672000:0.000000)0.992000:0.000000)0.308000:0.000000)0.876000:0.000000)1.000000:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)1.000000:0.000000)0.564000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.868000:0.000000)0.000000:0.000000,(PA38:0.000000,(PA40:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.336000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.140000:0.000000)0.120000:0.000000)0.088000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.516000:0.000000)0.040000:0.000000)0.124000:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000)0.884000:0.000000)0.820000:0.000000)0.092000:0.000000,(Dasypus_novemcinctus:0.000000,PA39:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.988000:0.000000)0.636000:0.000000)0.052000:0.000000)0.152000:0.000000)0.148000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.996000:0.000000)0.992000:0.000000)0.256000:0.000000,(Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.596000:0.000000)0.308000:0.000000)0.884000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.876000:0.000000)0.848000:0.000000)1.000000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“all characters” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: strict consensus of 10 MPTs - nodal values = "transfer bootstrap expectation" values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.896000:0.000000,((((PA3:0.000000,PA1:0.000000,(PA5:0.000000,(PA6:0.000000,(PA7:0.000000,(Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.440000:0.000000)0.832000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.744000:0.000000)0.606000:0.000000)0.591200:0.000000,(PA8:0.000000,(PA21:0.000000,(((Leptictis_dakotensis:0.000000,(Protungulatum_donnae:0.000000,(Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.368000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.272000:0.000000)0.309333:0.000000)0.387000:0.000000)0.416000:0.000000)0.355333:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)0.792000:0.000000,(Sinopa_rapax:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.670000:0.000000)1.000000:0.000000)0.397091:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.908000:0.000000)0.345538:0.000000)0.328857:0.000000,((PA9:0.000000,(PA10:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.988000:0.000000)0.610000:0.000000)0.878667:0.000000)0.748000:0.000000)0.665600:0.000000)0.644000:0.000000)0.625714:0.000000)0.590000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.928000:0.000000)0.618400:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.616000:0.000000)0.661667:0.000000)0.691692:0.000000)0.646857:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.688000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.776000:0.000000)0.990000:0.000000)1.000000:0.000000)0.998667:0.000000)1.000000:0.000000)0.548000:0.000000)0.339684:0.000000)0.295644:0.000000,(PA27:0.000000,PA33:0.000000,PA26:0.000000,(PA28:0.000000,(Ictidomys_tridecemlineatus:0.000000,Rattus_norvegicus:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.904000:0.000000,(Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.364000:0.000000,(Rhombomylus_turpanensis:0.000000,Tribosphenomys_minutus:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.672000:0.000000)0.992000:0.000000)0.703000:0.000000)0.987600:0.000000)1.000000:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)1.000000:0.000000)0.782000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.931130:0.000000)0.284000:0.000000,(PA38:0.000000,(PA40:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.336000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.140000:0.000000)0.248000:0.000000)0.300000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.516000:0.000000)0.320667:0.000000)0.344571:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000)0.884000:0.000000)0.844000:0.000000)0.391667:0.000000,(Dasypus_novemcinctus:0.000000,PA39:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.988000:0.000000)0.754667:0.000000)0.331294:0.000000)0.517500:0.000000)0.550571:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.996000:0.000000)0.993333:0.000000)0.680400:0.000000,(Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.596000:0.000000)0.526667:0.000000)0.928667:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.876000:0.000000)0.924000:0.000000)1.000000:0.000000)1.000000:0.000000));

[“all characters” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: 4 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((PA1,(PA3,((PA5,(PA6,((PA7,((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(Leptictis_dakotensis,Rhynchocyon_cirnei)))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))))),(PA38,(PA40,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Orycteropus_afer,(Manis_pentadactyla,Metacheiromys_marshi)))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((PA1,(PA3,((PA6,((PA7,((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(Leptictis_dakotensis,Rhynchocyon_cirnei)))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))),(PA5,((PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))),(PA40,(Orycteropus_afer,(Manis_pentadactyla,Metacheiromys_marshi)))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((PA1,(PA3,((PA5,(PA6,((PA7,((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(Leptictis_dakotensis,Rhynchocyon_cirnei)))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))))),((PA38,(PA39,(Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA40,(Orycteropus_afer,(Manis_pentadactyla,Metacheiromys_marshi))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((PA3,(PA1,((PA6,((PA7,((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(Leptictis_dakotensis,Rhynchocyon_cirnei)))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))),(PA5,(PA38,(PA40,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Orycteropus_afer,(Manis_pentadactyla,Metacheiromys_marshi))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));

[“all characters” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: strict consensus of 4 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.900000:0.000000,((((PA3:0.000000,PA1:0.000000,(PA5:0.000000,(PA6:0.000000,((PA7:0.000000,((Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.472000:0.000000)0.840000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.816000:0.000000)0.304000:0.000000)0.216000:0.000000,((((Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.268000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.356000:0.000000)0.092000:0.000000)0.268000:0.000000,(Protungulatum_donnae:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)1.000000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.696000:0.000000)1.000000:0.000000)0.056000:0.000000)0.036000:0.000000,(((Procavia_capensis:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.996000:0.000000)0.108000:0.000000)0.664000:0.000000)0.092000:0.000000)0.032000:0.000000)0.044000:0.000000)0.044000:0.000000)0.020000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.980000:0.000000)0.028000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.664000:0.000000)0.320000:0.000000)0.004000:0.000000,((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.036000:0.000000,(Moeritherium_lyonsi:0.000000,(Loxodonta_africana:0.000000,Trichechus_manatus:0.000000)0.132000:0.000000)0.336000:0.000000)0.004000:0.000000)0.164000:0.000000,((Rattus_norvegicus:0.000000,((Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.848000:0.000000,(Cocomys_lingchaensis:0.000000,Ictidomys_tridecemlineatus:0.000000)0.408000:0.000000)0.216000:0.000000)0.132000:0.000000,(Paramys_delicatus:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.848000:0.000000)1.000000:0.000000)0.456000:0.000000)0.292000:0.000000)0.048000:0.000000)1.000000:0.000000)0.052000:0.000000)0.088000:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.244000:0.000000)0.016000:0.000000)0.004000:0.000000)0.004000:0.000000,(PA26:0.000000,(((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.676000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.720000:0.000000)0.976000:0.000000)1.000000:0.000000)0.980000:0.000000)1.000000:0.000000)0.040000:0.000000,((Notharctus_tenebrosus:0.000000,(Lemur_catta:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.984000:0.000000)0.528000:0.000000)0.172000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.180000:0.000000)0.032000:0.000000)0.040000:0.000000)0.004000:0.000000)0.032000:0.000000,(PA40:0.000000,PA38:0.000000,(Dasypus_novemcinctus:0.000000,PA39:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.968000:0.000000)0.548000:0.000000,(Orycteropus_afer:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.964000:0.000000)0.192000:0.000000)0.172000:0.000000)0.328000:0.000000)0.328000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.996000:0.000000)0.992000:0.000000)0.464000:0.000000,(Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.692000:0.000000)0.440000:0.000000)0.896000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.880000:0.000000)0.812000:0.000000)1.000000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“all characters” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes aat the superordinal level and above included: strict consensus of 4 MPTs - nodal values = "transfer bootstrap expectation" values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.900000:0.000000,((((PA3:0.000000,PA1:0.000000,(PA5:0.000000,(PA6:0.000000,((PA7:0.000000,((Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.472000:0.000000)0.894000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.816000:0.000000)0.636000:0.000000)0.571200:0.000000,((((Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.268000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.356000:0.000000)0.357333:0.000000)0.434000:0.000000,(Protungulatum_donnae:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)1.000000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.898667:0.000000)1.000000:0.000000)0.811200:0.000000)0.432800:0.000000,(((Procavia_capensis:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.996000:0.000000)0.554000:0.000000)0.792000:0.000000)0.621000:0.000000)0.512800:0.000000)0.447333:0.000000)0.400000:0.000000)0.382000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.980000:0.000000)0.439200:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.664000:0.000000)0.522000:0.000000)0.499077:0.000000,((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.036000:0.000000,(Moeritherium_lyonsi:0.000000,(Loxodonta_africana:0.000000,Trichechus_manatus:0.000000)0.132000:0.000000)0.532000:0.000000)0.491000:0.000000)0.468222:0.000000,((Rattus_norvegicus:0.000000,((Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.848000:0.000000,(Cocomys_lingchaensis:0.000000,Ictidomys_tridecemlineatus:0.000000)0.408000:0.000000)0.541333:0.000000)0.514000:0.000000,(Paramys_delicatus:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.848000:0.000000)1.000000:0.000000)0.818667:0.000000)0.739000:0.000000)0.603200:0.000000)1.000000:0.000000)0.470897:0.000000)0.406400:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.244000:0.000000)0.389333:0.000000)0.362261:0.000000)0.353422:0.000000,(PA26:0.000000,(((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.676000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.720000:0.000000)0.982000:0.000000)1.000000:0.000000)0.996667:0.000000)1.000000:0.000000)0.786667:0.000000,((Notharctus_tenebrosus:0.000000,(Lemur_catta:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.984000:0.000000)0.756000:0.000000)0.724000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.726667:0.000000)0.500750:0.000000)0.515294:0.000000)0.382815:0.000000)0.397077:0.000000,(PA40:0.000000,PA38:0.000000,(Dasypus_novemcinctus:0.000000,PA39:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.968000:0.000000)0.692000:0.000000,(Orycteropus_afer:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.964000:0.000000)0.578000:0.000000)0.475000:0.000000)0.606000:0.000000)0.643429:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.996000:0.000000)0.993333:0.000000)0.758000:0.000000,(Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.692000:0.000000)0.650667:0.000000)0.933333:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.880000:0.000000)0.906000:0.000000)1.000000:0.000000)1.000000:0.000000)-1.#IND00:0.000000);

[“skeletal only" PAs corrected for the O'Leary et al. character ontology: 24 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((((PA5,((PA38,((Orycteropus_afer,(PA43,((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA6,((((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(PA11,((Sus_scrofa,(PA12,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15))),(Hippopotamus_amphibius,PA14))))),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA7,(((Erinaceus_europaeus,PA24),(Talpa_europaea,(Sorex_araneus,PA25))),(Solenodon_paradoxus,PA23)))),((PA28,((PA29,((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),(PA33,((PA35,(Tarsius_syrichta,(PA36,(PA37,(Homo_sapiens,Saimiri_sciureus))))),(Lemur_catta,Notharctus_tenebrosus)))),(PA26,(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,((PA38,((PA40,((Orycteropus_afer,(PA43,((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA5,((PA7,((PA8,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((Sus_scrofa,(PA12,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA23,(Solenodon_paradoxus,((Erinaceus_europaeus,PA24),(Sorex_araneus,(Talpa_europaea,PA25))))))),(PA6,((PA28,(((PA29,((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA33,((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(PA35,((Tarsius_syrichta,(PA36,(Homo_sapiens,(Saimiri_sciureus,PA37)))),(Lemur_catta,Notharctus_tenebrosus))))),(PA26,(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,((PA38,((PA40,((Orycteropus_afer,(PA43,((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA5,((PA7,((PA8,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((Sus_scrofa,(PA12,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA23,(Solenodon_paradoxus,(Erinaceus_europaeus,(PA24,(Talpa_europaea,(Sorex_araneus,PA25)))))))),(PA6,((PA28,(((PA29,(PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(PA33,((PA35,(PA36,(Tarsius_syrichta,(Homo_sapiens,(Saimiri_sciureus,PA37))))),(Lemur_catta,Notharctus_tenebrosus)))),(PA26,(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,(PA5,((((Orycteropus_afer,(PA43,((PA44,(Amblysomus_hottentotus,(Echinops_telfairi,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)))),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,(PA39,(Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA6,(((PA7,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,(Sus_scrofa,(PA12,(PA13,(Hippopotamus_amphibius,(PA14,((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15))))))))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA23,(Solenodon_paradoxus,((Erinaceus_europaeus,PA24),(Sorex_araneus,(Talpa_europaea,PA25)))))),((PA28,(((PA29,((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,(PA33,((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),((PA35,(Tarsius_syrichta,(PA36,(Saimiri_sciureus,(Homo_sapiens,PA37))))),(Lemur_catta,Notharctus_tenebrosus))))),(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,(PA5,((((Orycteropus_afer,(PA43,((PA44,(Echinops_telfairi,(Amblysomus_hottentotus,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)))),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA6,(((PA7,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,(Sus_scrofa,(PA12,(PA13,(Hippopotamus_amphibius,(PA14,((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))))))))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA23,(Solenodon_paradoxus,(Erinaceus_europaeus,(PA24,(Sorex_araneus,(Talpa_europaea,PA25))))))),((PA28,(((PA29,((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,(PA33,((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),((PA35,(Tarsius_syrichta,(PA36,(PA37,(Homo_sapiens,Saimiri_sciureus))))),(Lemur_catta,Notharctus_tenebrosus))))),(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,(((PA38,(PA40,((((PA44,(Echinops_telfairi,(Amblysomus_hottentotus,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(Orycteropus_afer,PA43)),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))),(PA5,((PA7,(((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((Sus_scrofa,(PA12,(PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15)))))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),((Erinaceus_europaeus,(PA24,(Talpa_europaea,(Sorex_araneus,PA25)))),(Solenodon_paradoxus,PA23)))),(PA6,((PA28,(((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(PA33,((PA36,(Tarsius_syrichta,(Homo_sapiens,(Saimiri_sciureus,PA37)))),(PA35,(Lemur_catta,Notharctus_tenebrosus))))),(PA26,(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA3,(PA1,(PA5,((((Orycteropus_afer,(PA43,(PA44,((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla))))),((PA6,(PA7,(((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((Sus_scrofa,(PA12,(PA13,(Hippopotamus_amphibius,(PA14,((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15)))))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),((Erinaceus_europaeus,(PA24,(Talpa_europaea,(Sorex_araneus,PA25)))),(Solenodon_paradoxus,PA23))))),((PA28,((PA29,(((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,((PA33,(Cynocephalus_volans,(Galeopterus_variegatus,PA34))),((PA35,(Tarsius_syrichta,(PA36,(Saimiri_sciureus,(Homo_sapiens,PA37))))),(Lemur_catta,Notharctus_tenebrosus)))),(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((((((PA40,(Orycteropus_afer,(PA43,((PA44,(Echinops_telfairi,(Amblysomus_hottentotus,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA5,((PA6,(((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12)),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),(PA7,(Solenodon_paradoxus,(PA23,(Erinaceus_europaeus,(PA24,(Talpa_europaea,(Sorex_araneus,PA25))))))))),((PA28,(((PA29,(PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(((PA33,(Galeopterus_variegatus,(Cynocephalus_volans,PA34))),(PA35,((Tarsius_syrichta,(PA36,(Saimiri_sciureus,(Homo_sapiens,PA37)))),(Lemur_catta,Notharctus_tenebrosus)))),(PA26,(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((((((PA40,(Orycteropus_afer,(PA43,((PA44,(Echinops_telfairi,(Amblysomus_hottentotus,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA5,((PA6,(((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12)),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),(PA7,(Solenodon_paradoxus,(PA23,(Erinaceus_europaeus,(PA24,(Sorex_araneus,(Talpa_europaea,PA25))))))))),((PA28,(((PA29,(PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(((PA33,(PA34,(Cynocephalus_volans,Galeopterus_variegatus))),((PA35,(Tarsius_syrichta,(PA36,(PA37,(Homo_sapiens,Saimiri_sciureus))))),(Lemur_catta,Notharctus_tenebrosus))),(PA26,(PA27,(PA32,(Ptilocercus_lowii,Tupaia_glis))))))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,(((PA40,((((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))),(Orycteropus_afer,PA43)),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA5,(PA6,((PA7,(((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(PA11,(((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12)),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),((Erinaceus_europaeus,(PA24,(Talpa_europaea,(Sorex_araneus,PA25)))),(Solenodon_paradoxus,PA23)))),((PA28,((((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),(PA33,((PA35,((PA37,(Homo_sapiens,Saimiri_sciureus)),(Tarsius_syrichta,PA36))),(Lemur_catta,Notharctus_tenebrosus))))),(PA27,(Tupaia_glis,(Ptilocercus_lowii,PA32)))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((((PA5,(PA38,(((PA40,(Orycteropus_afer,(PA43,(PA44,((PA45,(Amblysomus_hottentotus,Echinops_telfairi)),(Leptictis_dakotensis,Rhynchocyon_cirnei)))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(PA39,(Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)))))),((PA6,(((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA12,(Sus_scrofa,(PA13,(Hippopotamus_amphibius,(PA14,((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15)))))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),(PA7,(Solenodon_paradoxus,(PA23,((Erinaceus_europaeus,PA24),(Talpa_europaea,(Sorex_araneus,PA25)))))))),((PA28,((PA29,((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA26,(((PA33,(Galeopterus_variegatus,(Cynocephalus_volans,PA34))),(PA35,(((PA37,(Homo_sapiens,Saimiri_sciureus)),(Tarsius_syrichta,PA36)),(Lemur_catta,Notharctus_tenebrosus)))),(PA27,(PA32,(Ptilocercus_lowii,Tupaia_glis)))))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,((PA5,((PA38,(PA40,((Orycteropus_afer,(PA43,(PA44,((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla))))),((((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((Sus_scrofa,(PA12,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA7,(((Erinaceus_europaeus,PA24),(Sorex_araneus,(Talpa_europaea,PA25))),(Solenodon_paradoxus,PA23)))),(PA6,((PA28,(((PA29,((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),(PA33,((Tarsius_syrichta,(PA36,(Homo_sapiens,(Saimiri_sciureus,PA37)))),(PA35,(Lemur_catta,Notharctus_tenebrosus)))))),(PA27,(PA32,(Ptilocercus_lowii,Tupaia_glis))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,((PA5,(((((PA44,(Echinops_telfairi,(Amblysomus_hottentotus,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(Orycteropus_afer,PA43)),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA6,((PA7,(((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(PA11,(((PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15)))))),(Sus_scrofa,PA12)),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),(PA23,(Solenodon_paradoxus,(Erinaceus_europaeus,(PA24,(Talpa_europaea,(Sorex_araneus,PA25)))))))),((PA28,(((PA29,(PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA27,(PA26,((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(PA33,((PA35,((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36))),(Lemur_catta,Notharctus_tenebrosus)))))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((((PA5,(((((PA44,(Echinops_telfairi,(Amblysomus_hottentotus,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(Orycteropus_afer,PA43)),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA6,((PA7,(((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(PA11,(((PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15)))))),(Sus_scrofa,PA12)),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),(PA23,(Solenodon_paradoxus,(Erinaceus_europaeus,(PA24,(Talpa_europaea,(Sorex_araneus,PA25)))))))),((PA28,(((PA29,(PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA27,(PA26,((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(PA33,((PA35,((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36))),(Lemur_catta,Notharctus_tenebrosus)))))),(Ptilocercus_lowii,(Tupaia_glis,PA32))))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,((PA5,((PA38,(PA40,((((PA44,(Echinops_telfairi,(Amblysomus_hottentotus,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(Orycteropus_afer,PA43)),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),((((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(PA11,(((PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15)))))),(Sus_scrofa,PA12)),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA7,(PA23,(Solenodon_paradoxus,(Erinaceus_europaeus,(PA24,(Talpa_europaea,(Sorex_araneus,PA25)))))))),(PA6,((PA28,((PA29,((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(PA33,(((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36)),(PA35,(Lemur_catta,Notharctus_tenebrosus)))))),(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,(PA5,(((PA40,((Orycteropus_afer,(PA43,((PA44,(Amblysomus_hottentotus,(Echinops_telfairi,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA6,(((PA7,((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,(PA12,(Sus_scrofa,(PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))))))))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(((Erinaceus_europaeus,PA24),(PA25,(Sorex_araneus,Talpa_europaea))),(Solenodon_paradoxus,PA23))),((PA28,(((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA27,(PA26,((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(PA33,((Tarsius_syrichta,(PA36,(Homo_sapiens,(Saimiri_sciureus,PA37)))),(PA35,(Lemur_catta,Notharctus_tenebrosus))))))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,(PA5,(((PA38,(PA40,((PA43,(Orycteropus_afer,((PA44,(Amblysomus_hottentotus,(Echinops_telfairi,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))),(PA6,(((PA7,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,(Sus_scrofa,(PA12,(PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))))))))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),((Erinaceus_europaeus,(PA24,(PA25,(Sorex_araneus,Talpa_europaea)))),(Solenodon_paradoxus,PA23))),((PA28,(((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA27,(PA26,((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),(PA33,((PA35,((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36))),(Lemur_catta,Notharctus_tenebrosus)))))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,(PA5,(((PA38,(PA40,((((PA44,(Amblysomus_hottentotus,(Echinops_telfairi,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(Orycteropus_afer,PA43)),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))),(PA6,(((PA7,((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,(PA12,(Sus_scrofa,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15))),(Hippopotamus_amphibius,PA14)))))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(((Erinaceus_europaeus,PA24),(PA25,(Sorex_araneus,Talpa_europaea))),(Solenodon_paradoxus,PA23))),((PA28,(((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA27,(PA26,((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(PA33,(((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36)),(PA35,(Lemur_catta,Notharctus_tenebrosus))))))),(PA32,(Ptilocercus_lowii,Tupaia_glis)))))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,(PA2,PA1))),(((PA3,(PA5,((((PA40,(((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))),(Orycteropus_afer,PA43))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(PA38,(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA6,((((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,((PA13,(Hippopotamus_amphibius,(PA14,((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15)))))),(Sus_scrofa,PA12))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA7,(PA23,(Solenodon_paradoxus,(Erinaceus_europaeus,(PA24,(Talpa_europaea,(Sorex_araneus,PA25)))))))),((PA28,((PA29,(((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA27,(PA26,((PA33,(Galeopterus_variegatus,(Cynocephalus_volans,PA34))),(PA35,((Tarsius_syrichta,(PA36,(PA37,(Homo_sapiens,Saimiri_sciureus)))),(Lemur_catta,Notharctus_tenebrosus)))))),(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA2,(Tachyglossus_aculeatus,PA1))),(((PA3,(PA5,(((PA40,((((PA44,(Echinops_telfairi,(Amblysomus_hottentotus,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(Orycteropus_afer,PA43)),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla))))),((((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12)),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA7,(((Erinaceus_europaeus,PA24),(Talpa_europaea,(Sorex_araneus,PA25))),(Solenodon_paradoxus,PA23)))),(PA6,((PA28,((PA29,((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(PA33,((PA36,(Tarsius_syrichta,(Homo_sapiens,(Saimiri_sciureus,PA37)))),(PA35,(Lemur_catta,Notharctus_tenebrosus)))))),(PA27,(PA32,(Ptilocercus_lowii,Tupaia_glis)))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA2,(Tachyglossus_aculeatus,PA1))),(((PA3,((((PA43,(Orycteropus_afer,((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA5,(PA6,(((PA7,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(PA11,((PA12,(Sus_scrofa,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15))),(Hippopotamus_amphibius,PA14))))),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(Solenodon_paradoxus,(PA23,((Erinaceus_europaeus,PA24),(Sorex_araneus,(Talpa_europaea,PA25)))))),((PA28,(((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA27,((PA26,((PA33,(PA34,(Cynocephalus_volans,Galeopterus_variegatus))),((PA35,(PA36,(Tarsius_syrichta,(Saimiri_sciureus,(Homo_sapiens,PA37))))),(Lemur_catta,Notharctus_tenebrosus)))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),((((PA5,((PA38,((PA40,(((PA44,((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(Leptictis_dakotensis,Rhynchocyon_cirnei))),(Orycteropus_afer,PA43)),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA6,((((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,(Sus_scrofa,(PA12,(PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))))))))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),(PA7,(((Erinaceus_europaeus,PA24),(PA25,(Sorex_araneus,Talpa_europaea))),(Solenodon_paradoxus,PA23)))),((PA28,((PA29,(((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(PA33,(((Homo_sapiens,(Saimiri_sciureus,PA37)),(Tarsius_syrichta,PA36)),(PA35,(Lemur_catta,Notharctus_tenebrosus)))))),(PA27,(Tupaia_glis,(Ptilocercus_lowii,PA32))))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,((PA5,((PA40,(((PA44,((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(Leptictis_dakotensis,Rhynchocyon_cirnei))),(Orycteropus_afer,PA43)),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla)))))),(((PA8,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA7,(Solenodon_paradoxus,(PA23,((Erinaceus_europaeus,PA24),(Talpa_europaea,(Sorex_araneus,PA25))))))),(PA6,((PA28,(((PA29,(PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA27,(PA26,((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(PA33,((PA35,((Homo_sapiens,(Saimiri_sciureus,PA37)),(Tarsius_syrichta,PA36))),(Lemur_catta,Notharctus_tenebrosus)))))),(Tupaia_glis,(Ptilocercus_lowii,PA32)))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA2,(Tachyglossus_aculeatus,PA1))),(((PA3,(PA5,((PA38,(((((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))),(Orycteropus_afer,PA43)),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA39,(Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla))))),((PA6,(((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,(Sus_scrofa,(PA12,(PA13,(Hippopotamus_amphibius,(PA14,((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))))))))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),(PA7,(Solenodon_paradoxus,(PA23,((Erinaceus_europaeus,PA24),(PA25,(Sorex_araneus,Talpa_europaea)))))))),((PA28,((PA29,((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA26,(((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),(PA33,((Tarsius_syrichta,(PA36,(Saimiri_sciureus,(Homo_sapiens,PA37)))),(PA35,(Lemur_catta,Notharctus_tenebrosus))))),(PA27,(PA32,(Ptilocercus_lowii,Tupaia_glis)))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));

[“skeletal only” PAs corrected for the O'Leary et al. character ontology: strict consensus of 24 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000,PA3:0.000000,PA2:0.000000,PA1:0.000000,(PA5:0.000000,(PA38:0.000000,(PA40:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(PA44:0.000000,(Amblysomus_hottentotus:0.000000,Echinops_telfairi:0.000000,PA45:0.000000)0.460000:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.024000:0.000000)0.016000:0.000000)0.008000:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.244000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.100000:0.000000)0.088000:0.000000)0.048000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.308000:0.000000)0.024000:0.000000)0.084000:0.000000)0.000000:0.000000,(Dasypus_novemcinctus:0.000000,PA39:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.988000:0.000000)0.616000:0.000000)0.000000:0.000000,(PA6:0.000000,(PA7:0.000000,(PA8:0.000000,(((((Apheliscus_insidiosus:0.000000,Protungulatum_donnae:0.000000)0.016000:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.312000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.252000:0.000000)0.052000:0.000000)0.164000:0.000000,(Rodhocetus_balochistanensis:0.000000,(Archaeotherium_mortoni:0.000000,Mesonyx_obtusidens:0.000000)0.044000:0.000000)0.080000:0.000000)0.000000:0.000000,((PA10:0.000000,(PA11:0.000000,(Sus_scrofa:0.000000,PA12:0.000000,(PA13:0.000000,(Hippopotamus_amphibius:0.000000,PA14:0.000000,((Artiocetus_clavis:0.000000,Basilosaurus_cetoides:0.000000)0.232000:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000,PA15:0.000000)0.996000:0.000000)0.156000:0.000000)0.148000:0.000000)0.004000:0.000000)0.004000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.044000:0.000000)0.016000:0.000000)0.000000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.360000:0.000000)0.012000:0.000000)0.004000:0.000000,(PA9:0.000000,(((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,(((Myotis_lucifugus:0.000000,PA17:0.000000)0.216000:0.000000,(Pteronotus_parnellii:0.000000,PA18:0.000000)0.104000:0.000000)0.424000:0.000000,((Nycteris_thebaica:0.000000,PA19:0.000000)0.052000:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.428000:0.000000)0.756000:0.000000)0.600000:0.000000)0.424000:0.000000,(PA16:0.000000,(Pteropus_giganteus:0.000000,PA20:0.000000)0.428000:0.000000)0.092000:0.000000)1.000000:0.000000)0.092000:0.000000)0.000000:0.000000,(PA21:0.000000,(((Canis_lupus:0.000000,Felis_silvestris:0.000000,PA22:0.000000)0.760000:0.000000,(Sinopa_rapax:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.924000:0.000000)0.948000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.956000:0.000000)0.064000:0.000000)0.164000:0.000000)0.000000:0.000000,(Solenodon_paradoxus:0.000000,PA23:0.000000,(Erinaceus_europaeus:0.000000,PA24:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000,PA25:0.000000)0.816000:0.000000)0.160000:0.000000)0.156000:0.000000)0.000000:0.000000,((PA28:0.000000,((PA29:0.000000,(Ictidomys_tridecemlineatus:0.000000,PA31:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.192000:0.000000)0.116000:0.000000,(Rattus_norvegicus:0.000000,PA30:0.000000)0.188000:0.000000)0.220000:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.596000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.756000:0.000000)0.980000:0.000000)0.380000:0.000000)0.340000:0.000000)0.312000:0.000000)0.524000:0.000000)1.000000:0.000000,(PA27:0.000000,PA26:0.000000,(PA33:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000,PA34:0.000000)1.000000:0.000000,(PA35:0.000000,(Tarsius_syrichta:0.000000,PA36:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000,PA37:0.000000)1.000000:0.000000)0.512000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000)0.356000:0.000000)1.000000:0.000000)0.748000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000,PA32:0.000000)1.000000:0.000000)0.464000:0.000000)0.728000:0.000000)0.000000:0.000000)0.112000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.984000:0.000000)0.980000:0.000000,((Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000)0.472000:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.344000:0.000000)0.600000:0.000000)0.600000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.972000:0.000000)1.000000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“skeletal only” PAs corrected for the O'Leary et al. character ontology: strict consensus of 24 MPTs - nodal values = "transfer bootstrap expectation" values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000,PA3:0.000000,PA2:0.000000,PA1:0.000000,(PA5:0.000000,(PA38:0.000000,(PA40:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(PA44:0.000000,(Amblysomus_hottentotus:0.000000,Echinops_telfairi:0.000000,PA45:0.000000)0.492000:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.024000:0.000000)0.244800:0.000000)0.183429:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.244000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.100000:0.000000)0.194000:0.000000)0.236000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.308000:0.000000)0.258000:0.000000)0.276571:0.000000)0.161500:0.000000,(Dasypus_novemcinctus:0.000000,PA39:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.988000:0.000000)0.742667:0.000000)0.161524:0.000000,(PA6:0.000000,(PA7:0.000000,(PA8:0.000000,(((((Apheliscus_insidiosus:0.000000,Protungulatum_donnae:0.000000)0.016000:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.312000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.252000:0.000000)0.252000:0.000000)0.315200:0.000000,(Rodhocetus_balochistanensis:0.000000,(Archaeotherium_mortoni:0.000000,Mesonyx_obtusidens:0.000000)0.044000:0.000000)0.128000:0.000000)0.215500:0.000000,((PA10:0.000000,(PA11:0.000000,(Sus_scrofa:0.000000,PA12:0.000000,(PA13:0.000000,(Hippopotamus_amphibius:0.000000,PA14:0.000000,((Artiocetus_clavis:0.000000,Basilosaurus_cetoides:0.000000)0.232000:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000,PA15:0.000000)0.996000:0.000000)0.610000:0.000000)0.492667:0.000000)0.424000:0.000000)0.339111:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.044000:0.000000)0.378333:0.000000)0.393538:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.360000:0.000000)0.432800:0.000000)0.376500:0.000000,(PA9:0.000000,(((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,(((Myotis_lucifugus:0.000000,PA17:0.000000)0.216000:0.000000,(Pteronotus_parnellii:0.000000,PA18:0.000000)0.104000:0.000000)0.432000:0.000000,((Nycteris_thebaica:0.000000,PA19:0.000000)0.052000:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.428000:0.000000)0.796000:0.000000)0.772000:0.000000)0.834667:0.000000,(PA16:0.000000,(Pteropus_giganteus:0.000000,PA20:0.000000)0.428000:0.000000)0.260000:0.000000)1.000000:0.000000)0.930154:0.000000)0.386421:0.000000,(PA21:0.000000,(((Canis_lupus:0.000000,Felis_silvestris:0.000000,PA22:0.000000)0.760000:0.000000,(Sinopa_rapax:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.962000:0.000000)0.968800:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.956000:0.000000)0.724571:0.000000)0.687000:0.000000)0.308500:0.000000,(Solenodon_paradoxus:0.000000,PA23:0.000000,(Erinaceus_europaeus:0.000000,PA24:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000,PA25:0.000000)0.822000:0.000000)0.530000:0.000000)0.430667:0.000000)0.280857:0.000000,((PA28:0.000000,((PA29:0.000000,(Ictidomys_tridecemlineatus:0.000000,PA31:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.192000:0.000000)0.157333:0.000000,(Rattus_norvegicus:0.000000,PA30:0.000000)0.188000:0.000000)0.298000:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.596000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.756000:0.000000)0.980000:0.000000)0.782667:0.000000)0.731000:0.000000)0.636000:0.000000)0.963385:0.000000)1.000000:0.000000,(PA27:0.000000,PA26:0.000000,(PA33:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000,PA34:0.000000)1.000000:0.000000,(PA35:0.000000,(Tarsius_syrichta:0.000000,PA36:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000,PA37:0.000000)1.000000:0.000000)0.815000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000)0.356000:0.000000)1.000000:0.000000)0.909455:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000,PA32:0.000000)1.000000:0.000000)0.774750:0.000000)0.859613:0.000000)0.183000:0.000000)0.409412:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.984000:0.000000)0.981333:0.000000,((Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000)0.472000:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.344000:0.000000)0.694667:0.000000)0.804000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.972000:0.000000)1.000000:0.000000)1.000000:0.000000));

[“skeletal only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: 8 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),(((((PA5,((((PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus)))),(PA8,((PA21,(((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))),(Manis_pentadactyla,Metacheiromys_marshi))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))),(PA6,((PA26,(PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))))),((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))),((PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(Orycteropus_afer,(PA43,(Rhynchocyon_cirnei,PA44))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((PA3,(PA1,((((PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus)))),((PA8,(PA21,(((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))),(Manis_pentadactyla,Metacheiromys_marshi)))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))))),(PA6,(PA26,((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))))),(PA5,((PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA43,(Orycteropus_afer,(Rhynchocyon_cirnei,PA44))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((((Solenodon_paradoxus,(Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea))),(Echinops_telfairi,Erinaceus_europaeus)),(Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))))),(((PA6,(PA7,(PA8,(((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),(PA21,(((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))))),((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA26,(PA33,(((Cynocephalus_volans,Galeopterus_variegatus),(Ptilocercus_lowii,Tupaia_glis)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus))))))),(PA5,((PA38,(PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(Orycteropus_afer,(PA43,(Rhynchocyon_cirnei,PA44)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))))),(PA3,PA1)),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((PA5,((((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(((Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))),(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus)))),(Leptictis_dakotensis,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei))))),(PA7,(PA8,((PA9,((PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))),(PA21,(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA6,(PA26,((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))))),((PA38,((PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),((Orycteropus_afer,PA43),(Rhynchocyon_cirnei,PA44)))),(PA39,(Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((((Solenodon_paradoxus,(Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea))),(Echinops_telfairi,Erinaceus_europaeus)),(Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))))),((PA5,(PA6,((PA7,(PA8,(((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),(PA21,(((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA26,(PA33,(((Cynocephalus_volans,Galeopterus_variegatus),(Ptilocercus_lowii,Tupaia_glis)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus))))))))),((PA38,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA40,(Orycteropus_afer,(PA43,(Rhynchocyon_cirnei,PA44)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((((Solenodon_paradoxus,(Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea))),(Echinops_telfairi,Erinaceus_europaeus)),(Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))))),(((PA6,(PA7,(((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),(PA8,(PA21,(((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))))),((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA26,(PA33,(((Cynocephalus_volans,Galeopterus_variegatus),(Ptilocercus_lowii,Tupaia_glis)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus))))))),(PA5,(PA38,(((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA40,((Orycteropus_afer,PA43),(Rhynchocyon_cirnei,PA44)))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))))))),(PA3,PA1)),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((PA5,((PA7,(((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(((Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))),(Leptictis_dakotensis,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),((PA8,(PA9,((PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))))),(PA21,(Manis_pentadactyla,Metacheiromys_marshi))))),(PA6,(PA26,((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))))),(((PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(Orycteropus_afer,(PA43,(Rhynchocyon_cirnei,PA44)))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((((Solenodon_paradoxus,(Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea))),(Echinops_telfairi,Erinaceus_europaeus)),(Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))))),((PA5,((PA6,(PA7,((PA8,((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))),(PA21,(((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA26,(PA33,(((Cynocephalus_volans,Galeopterus_variegatus),(Ptilocercus_lowii,Tupaia_glis)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus)))))))),((PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA43,(Orycteropus_afer,(Rhynchocyon_cirnei,PA44))))),(PA38,(PA39,(Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));

[“skeletal only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: strict consensus of 8 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.924000:0.000000,((Eomaia_scansoria:0.000000,Leptictis_dakotensis:0.000000,Maelestes_gobiensis:0.000000,PA21:0.000000,PA8:0.000000,PA7:0.000000,PA6:0.000000,PA5:0.000000,PA3:0.000000,PA1:0.000000,(Solenodon_paradoxus:0.000000,(Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.460000:0.000000)0.792000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.740000:0.000000)0.248000:0.000000,(Protungulatum_donnae:0.000000,(Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.368000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.264000:0.000000)0.080000:0.000000)0.152000:0.000000)0.208000:0.000000,(PA9:0.000000,(PA10:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.992000:0.000000)0.192000:0.000000)0.776000:0.000000)0.268000:0.000000)0.168000:0.000000)0.208000:0.000000)0.240000:0.000000)0.212000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.924000:0.000000)0.176000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.512000:0.000000)0.076000:0.000000)0.516000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.684000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.752000:0.000000)0.988000:0.000000)1.000000:0.000000)0.992000:0.000000)1.000000:0.000000)0.072000:0.000000,(Sinopa_rapax:0.000000,(Canis_lupus:0.000000,Felis_silvestris:0.000000)0.812000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)1.000000:0.000000,(PA38:0.000000,(PA40:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.240000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.116000:0.000000)0.116000:0.000000)0.060000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.464000:0.000000)0.012000:0.000000)0.060000:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000)0.884000:0.000000)0.680000:0.000000)0.032000:0.000000,(Dasypus_novemcinctus:0.000000,PA39:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.988000:0.000000)0.640000:0.000000)0.008000:0.000000,(PA26:0.000000,(PA27:0.000000,(PA28:0.000000,((Ictidomys_tridecemlineatus:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.900000:0.000000)0.284000:0.000000,(Rattus_norvegicus:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.344000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.664000:0.000000)0.996000:0.000000)0.316000:0.000000)0.320000:0.000000)0.152000:0.000000)0.120000:0.000000)0.816000:0.000000)1.000000:0.000000)0.112000:0.000000,(PA33:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.996000:0.000000)0.512000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.128000:0.000000)0.728000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.992000:0.000000)0.988000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.904000:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.604000:0.000000)0.908000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.860000:0.000000)0.860000:0.000000)1.000000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“skeletal only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: strict consensus of 8 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.924000:0.000000,((Eomaia_scansoria:0.000000,Leptictis_dakotensis:0.000000,Maelestes_gobiensis:0.000000,PA21:0.000000,PA8:0.000000,PA7:0.000000,PA6:0.000000,PA5:0.000000,PA3:0.000000,PA1:0.000000,(Solenodon_paradoxus:0.000000,(Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.460000:0.000000)0.844000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.740000:0.000000)0.576800:0.000000,(Protungulatum_donnae:0.000000,(Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.368000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.264000:0.000000)0.337333:0.000000)0.422000:0.000000)0.426400:0.000000,(PA9:0.000000,(PA10:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.992000:0.000000)0.594000:0.000000)0.873333:0.000000)0.727000:0.000000)0.636800:0.000000)0.601333:0.000000)0.571429:0.000000)0.531500:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.924000:0.000000)0.554800:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.512000:0.000000)0.596667:0.000000)0.627385:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.684000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.752000:0.000000)0.992000:0.000000)1.000000:0.000000)0.998667:0.000000)1.000000:0.000000)0.493091:0.000000,(Sinopa_rapax:0.000000,(Canis_lupus:0.000000,Felis_silvestris:0.000000)0.812000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)1.000000:0.000000,(PA38:0.000000,(PA40:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.240000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.116000:0.000000)0.238000:0.000000)0.266000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.464000:0.000000)0.254667:0.000000)0.262286:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000)0.884000:0.000000)0.749333:0.000000)0.290000:0.000000,(Dasypus_novemcinctus:0.000000,PA39:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.988000:0.000000)0.757333:0.000000)0.240706:0.000000,(PA26:0.000000,(PA27:0.000000,(PA28:0.000000,((Ictidomys_tridecemlineatus:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.900000:0.000000)0.612000:0.000000,(Rattus_norvegicus:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.344000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.664000:0.000000)0.996000:0.000000)0.769333:0.000000)0.707000:0.000000)0.542667:0.000000)0.617143:0.000000)0.981600:0.000000)1.000000:0.000000)0.926000:0.000000,(PA33:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.996000:0.000000)0.754000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.611556:0.000000)0.858087:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.992000:0.000000)0.990667:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.904000:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.604000:0.000000)0.946000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.860000:0.000000)0.930000:0.000000)1.000000:0.000000)1.000000:0.000000)-1.#IND00:0.000000);

[“skeletal only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: 2 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((PA1,(PA3,((PA6,((PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus)))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))),(PA5,(((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(PA38,(PA40,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Orycteropus_afer,(Manis_pentadactyla,Metacheiromys_marshi)))))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),(((((PA5,((PA6,((PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus)))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))),(((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(PA38,(PA40,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Orycteropus_afer,(Manis_pentadactyla,Metacheiromys_marshi)))))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));

[“skeletal only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: strict consensus of 2 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.904000:0.000000,((((PA3:0.000000,PA1:0.000000,(PA5:0.000000,(PA6:0.000000,((PA7:0.000000,(Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.480000:0.000000)0.812000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.820000:0.000000)0.240000:0.000000)0.232000:0.000000)0.004000:0.000000,(PA26:0.000000,(((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.652000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.740000:0.000000)0.972000:0.000000)1.000000:0.000000)0.988000:0.000000)1.000000:0.000000)0.044000:0.000000,((Notharctus_tenebrosus:0.000000,(Lemur_catta:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.984000:0.000000)0.480000:0.000000)0.160000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.188000:0.000000)0.032000:0.000000)0.044000:0.000000)0.000000:0.000000)0.008000:0.000000,(((((Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.308000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.360000:0.000000)0.080000:0.000000)0.280000:0.000000,(Protungulatum_donnae:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)0.984000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.672000:0.000000)1.000000:0.000000)0.068000:0.000000)0.040000:0.000000,(((Procavia_capensis:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.992000:0.000000)0.112000:0.000000)0.616000:0.000000)0.096000:0.000000)0.028000:0.000000)0.056000:0.000000)0.048000:0.000000)0.028000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.984000:0.000000)0.032000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.612000:0.000000)0.284000:0.000000)0.012000:0.000000,((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.032000:0.000000,(Moeritherium_lyonsi:0.000000,(Loxodonta_africana:0.000000,Trichechus_manatus:0.000000)0.160000:0.000000)0.332000:0.000000)0.016000:0.000000)0.200000:0.000000,((Rattus_norvegicus:0.000000,((Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.852000:0.000000,(Cocomys_lingchaensis:0.000000,Ictidomys_tridecemlineatus:0.000000)0.380000:0.000000)0.216000:0.000000)0.124000:0.000000,(Paramys_delicatus:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.852000:0.000000)1.000000:0.000000)0.440000:0.000000)0.284000:0.000000)0.044000:0.000000)1.000000:0.000000)0.044000:0.000000)0.104000:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.268000:0.000000)0.024000:0.000000,(PA38:0.000000,(PA40:0.000000,(((Dasypus_novemcinctus:0.000000,PA39:0.000000)0.212000:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.984000:0.000000)0.580000:0.000000,(Orycteropus_afer:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.976000:0.000000)0.212000:0.000000)0.096000:0.000000)0.008000:0.000000)0.188000:0.000000)0.008000:0.000000)0.316000:0.000000)0.316000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.992000:0.000000)0.988000:0.000000)0.480000:0.000000,(Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.716000:0.000000)0.456000:0.000000)0.900000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.888000:0.000000)0.804000:0.000000)1.000000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“skeletal only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: strict consensus of 2 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.904000:0.000000,((((PA3:0.000000,PA1:0.000000,(PA5:0.000000,(PA6:0.000000,((PA7:0.000000,(Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.480000:0.000000)0.874000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.820000:0.000000)0.612000:0.000000)0.567200:0.000000)0.473333:0.000000,(PA26:0.000000,(((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.652000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.740000:0.000000)0.980000:0.000000)1.000000:0.000000)0.998000:0.000000)1.000000:0.000000)0.787556:0.000000,((Notharctus_tenebrosus:0.000000,(Lemur_catta:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.984000:0.000000)0.732000:0.000000)0.720000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.729333:0.000000)0.496500:0.000000)0.511529:0.000000)0.375667:0.000000)0.361280:0.000000,(((((Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.308000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.360000:0.000000)0.357333:0.000000)0.433000:0.000000,(Protungulatum_donnae:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)0.984000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.890667:0.000000)1.000000:0.000000)0.813600:0.000000)0.434000:0.000000,(((Procavia_capensis:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.992000:0.000000)0.556000:0.000000)0.766667:0.000000)0.601000:0.000000)0.495200:0.000000)0.436667:0.000000)0.393143:0.000000)0.374500:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.984000:0.000000)0.412000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.612000:0.000000)0.491333:0.000000)0.473846:0.000000,((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.032000:0.000000,(Moeritherium_lyonsi:0.000000,(Loxodonta_africana:0.000000,Trichechus_manatus:0.000000)0.160000:0.000000)0.534000:0.000000)0.500000:0.000000)0.463111:0.000000,((Rattus_norvegicus:0.000000,((Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.852000:0.000000,(Cocomys_lingchaensis:0.000000,Ictidomys_tridecemlineatus:0.000000)0.380000:0.000000)0.528000:0.000000)0.524000:0.000000,(Paramys_delicatus:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.852000:0.000000)1.000000:0.000000)0.813333:0.000000)0.729000:0.000000)0.594400:0.000000)1.000000:0.000000)0.471448:0.000000)0.421300:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.268000:0.000000)0.405524:0.000000,(PA38:0.000000,(PA40:0.000000,(((Dasypus_novemcinctus:0.000000,PA39:0.000000)0.212000:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.984000:0.000000)0.716000:0.000000,(Orycteropus_afer:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.976000:0.000000)0.594000:0.000000)0.536000:0.000000)0.490286:0.000000)0.480000:0.000000)0.347535:0.000000)0.610500:0.000000)0.649143:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.992000:0.000000)0.990667:0.000000)0.766800:0.000000,(Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.716000:0.000000)0.654667:0.000000)0.936000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.888000:0.000000)0.902000:0.000000)1.000000:0.000000)1.000000:0.000000)-1.#IND00:0.000000);

[“craniodental only" PAs corrected for the O'Leary et al. character ontology: 12 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,(PA5,((PA38,((PA40,(PA43,(PA44,(Rhynchocyon_cirnei,((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(Leptictis_dakotensis,(Erinaceus_europaeus,((Solenodon_paradoxus,PA23),(PA24,(Talpa_europaea,(Sorex_araneus,PA25))))))))))),((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(Orycteropus_afer,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA6,((PA7,(PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),((Archaeotherium_mortoni,Mesonyx_obtusidens),(Rodhocetus_balochistanensis,((PA10,(PA11,(((PA13,(Hippopotamus_amphibius,(PA14,((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15)))))),(Sus_scrofa,PA12)),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))))))),((PA27,(PA28,(((PA29,(Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus)))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(PA26,((PA33,((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),(Ptilocercus_lowii,(Tupaia_glis,PA32)))),((PA36,(Tarsius_syrichta,(Saimiri_sciureus,(Homo_sapiens,PA37)))),(PA35,(Lemur_catta,Notharctus_tenebrosus)))))))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,(PA5,((PA38,((PA40,(PA43,(PA44,(Rhynchocyon_cirnei,((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(Leptictis_dakotensis,(Erinaceus_europaeus,((Solenodon_paradoxus,PA23),(PA24,(Sorex_araneus,(Talpa_europaea,PA25))))))))))),((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(Orycteropus_afer,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA6,((PA7,(PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),((Archaeotherium_mortoni,Mesonyx_obtusidens),(Rodhocetus_balochistanensis,((PA10,(PA11,(((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12)),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))))))),((PA27,(PA28,(((PA29,((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31))),(Rattus_norvegicus,PA30)),((Tribosphenomys_minutus,(Cocomys_lingchaensis,Paramys_delicatus)),(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA26,((PA33,((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(Ptilocercus_lowii,(Tupaia_glis,PA32)))),((PA36,(Tarsius_syrichta,(Saimiri_sciureus,(Homo_sapiens,PA37)))),(PA35,(Lemur_catta,Notharctus_tenebrosus)))))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((((PA5,(PA38,((PA43,(((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(Leptictis_dakotensis,(Erinaceus_europaeus,((PA24,(Solenodon_paradoxus,PA23)),(Sorex_araneus,(Talpa_europaea,PA25)))))),(Rhynchocyon_cirnei,PA44))),((PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(Orycteropus_afer,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))))),((PA7,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),((Archaeotherium_mortoni,Mesonyx_obtusidens),(Rodhocetus_balochistanensis,((PA10,((Sus_scrofa,(PA12,(PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15)))))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,((PA22,(Canis_lupus,Felis_silvestris)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))))))),(PA6,((PA26,(PA27,(PA28,((PA29,((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA33,(((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(Ptilocercus_lowii,(Tupaia_glis,PA32))),((PA35,((PA37,(Homo_sapiens,Saimiri_sciureus)),(Tarsius_syrichta,PA36))),(Lemur_catta,Notharctus_tenebrosus)))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,((PA38,(PA40,((PA43,(((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(Leptictis_dakotensis,(Erinaceus_europaeus,((Solenodon_paradoxus,PA23),(PA24,(Sorex_araneus,(Talpa_europaea,PA25))))))),(Rhynchocyon_cirnei,PA44))),((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(Orycteropus_afer,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))))),(PA5,(PA6,((PA7,((PA8,((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),((Archaeotherium_mortoni,Mesonyx_obtusidens),(Rodhocetus_balochistanensis,((PA10,(((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12)),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,((PA22,(Canis_lupus,Felis_silvestris)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))))),(PA26,((PA27,(PA28,((PA29,((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(PA33,(((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(Ptilocercus_lowii,(Tupaia_glis,PA32))),((PA35,(PA36,(Tarsius_syrichta,(Saimiri_sciureus,(Homo_sapiens,PA37))))),(Lemur_catta,Notharctus_tenebrosus))))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((((PA5,(PA38,((PA43,(Rhynchocyon_cirnei,(PA44,((PA45,(Amblysomus_hottentotus,Echinops_telfairi)),(Leptictis_dakotensis,(Erinaceus_europaeus,((PA24,(Solenodon_paradoxus,PA23)),(Sorex_araneus,(Talpa_europaea,PA25))))))))),(PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(Orycteropus_afer,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))))),(PA6,((PA7,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),((Archaeotherium_mortoni,Mesonyx_obtusidens),(Rodhocetus_balochistanensis,((PA10,(PA11,(((PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15)))))),(Sus_scrofa,PA12)),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,((PA22,(Canis_lupus,Felis_silvestris)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))))))),((PA26,(PA27,(PA28,((PA29,((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA33,(((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(Ptilocercus_lowii,(Tupaia_glis,PA32))),((PA35,(PA36,(Tarsius_syrichta,(Homo_sapiens,(Saimiri_sciureus,PA37))))),(Lemur_catta,Notharctus_tenebrosus)))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,((PA5,(PA38,((PA43,(Rhynchocyon_cirnei,(PA44,((PA45,(Amblysomus_hottentotus,Echinops_telfairi)),(Leptictis_dakotensis,(Erinaceus_europaeus,((PA24,(Solenodon_paradoxus,PA23)),(Sorex_araneus,(Talpa_europaea,PA25))))))))),(PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(Orycteropus_afer,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))))),(PA6,((PA7,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),((Archaeotherium_mortoni,Mesonyx_obtusidens),(Rodhocetus_balochistanensis,((PA10,(PA11,(((PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15)))))),(Sus_scrofa,PA12)),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,((PA22,(Canis_lupus,Felis_silvestris)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))))))),((PA26,(PA27,(PA28,((PA29,((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA33,(((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(Ptilocercus_lowii,(Tupaia_glis,PA32))),((PA35,(PA36,(Tarsius_syrichta,(Homo_sapiens,(Saimiri_sciureus,PA37))))),(Lemur_catta,Notharctus_tenebrosus))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,((PA38,((PA43,(((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(Leptictis_dakotensis,(Erinaceus_europaeus,(PA24,((Solenodon_paradoxus,PA23),(Sorex_araneus,(Talpa_europaea,PA25))))))),(Rhynchocyon_cirnei,PA44))),((PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(Orycteropus_afer,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA5,((PA6,(PA7,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),((Archaeotherium_mortoni,Mesonyx_obtusidens),(Rodhocetus_balochistanensis,((PA10,((Sus_scrofa,(PA12,(PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15)))))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,((PA22,(Canis_lupus,Felis_silvestris)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))))))),((PA26,(PA27,(PA28,(((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(Ptilocercus_lowii,(Tupaia_glis,PA32))),(PA33,(PA35,(((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36)),(Lemur_catta,Notharctus_tenebrosus)))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA3,(PA1,((PA38,((PA43,(((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(Leptictis_dakotensis,(Erinaceus_europaeus,((PA24,(Solenodon_paradoxus,PA23)),(Sorex_araneus,(Talpa_europaea,PA25)))))),(Rhynchocyon_cirnei,PA44))),((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA40,(Orycteropus_afer,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))))),(PA5,((PA6,(PA7,(PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),((Archaeotherium_mortoni,Mesonyx_obtusidens),(Rodhocetus_balochistanensis,((PA10,(PA11,(((PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15)))))),(Sus_scrofa,PA12)),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,((PA22,(Canis_lupus,Felis_silvestris)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))))))),((PA26,(PA27,(PA28,((PA29,((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Tribosphenomys_minutus,(Cocomys_lingchaensis,Paramys_delicatus)),(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(Ptilocercus_lowii,(Tupaia_glis,PA32))),(PA33,((PA35,((PA37,(Homo_sapiens,Saimiri_sciureus)),(Tarsius_syrichta,PA36))),(Lemur_catta,Notharctus_tenebrosus)))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,(PA5,((PA38,((PA40,(PA43,(Rhynchocyon_cirnei,(PA44,((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(Leptictis_dakotensis,(Erinaceus_europaeus,(PA24,((Solenodon_paradoxus,PA23),(Sorex_araneus,(Talpa_europaea,PA25))))))))))),((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(Orycteropus_afer,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))),((PA6,(PA7,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),((Archaeotherium_mortoni,Mesonyx_obtusidens),(Rodhocetus_balochistanensis,((PA10,(PA11,(((PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus)))))),(Sus_scrofa,PA12)),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,((PA22,(Canis_lupus,Felis_silvestris)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))))))),(PA26,((PA27,(PA28,((PA29,(((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(Rattus_norvegicus,PA30))),((Tribosphenomys_minutus,(Cocomys_lingchaensis,Paramys_delicatus)),(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA33,((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(Ptilocercus_lowii,(Tupaia_glis,PA32)))),((PA35,(PA36,(Tarsius_syrichta,(PA37,(Homo_sapiens,Saimiri_sciureus))))),(Lemur_catta,Notharctus_tenebrosus))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((((PA5,(PA38,((PA43,(((PA45,(Amblysomus_hottentotus,Echinops_telfairi)),(Leptictis_dakotensis,(Erinaceus_europaeus,(PA24,((Solenodon_paradoxus,PA23),(Talpa_europaea,(Sorex_araneus,PA25))))))),(Rhynchocyon_cirnei,PA44))),(PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(Orycteropus_afer,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))))),((PA7,((PA8,((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),((Archaeotherium_mortoni,Mesonyx_obtusidens),(Rodhocetus_balochistanensis,((PA10,(((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12)),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))))),(PA6,((PA26,(PA27,(PA28,(((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(PA29,(Rattus_norvegicus,PA30))),((Tribosphenomys_minutus,(Cocomys_lingchaensis,Paramys_delicatus)),(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),((PA33,((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(Ptilocercus_lowii,(Tupaia_glis,PA32)))),(PA35,(((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36)),(Lemur_catta,Notharctus_tenebrosus)))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,((PA5,(PA38,((PA43,(Rhynchocyon_cirnei,(PA44,((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(Leptictis_dakotensis,(Erinaceus_europaeus,((Solenodon_paradoxus,PA23),(PA24,(Sorex_araneus,(Talpa_europaea,PA25)))))))))),(PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(Orycteropus_afer,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))))),((PA7,(PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),((Archaeotherium_mortoni,Mesonyx_obtusidens),(Rodhocetus_balochistanensis,((PA10,(((PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15)))))),(Sus_scrofa,PA12)),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))))))),(PA6,((PA26,(PA27,(PA28,(((PA29,(PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)))),(Rattus_norvegicus,PA30)),((Tribosphenomys_minutus,(Cocomys_lingchaensis,Paramys_delicatus)),(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(Ptilocercus_lowii,(Tupaia_glis,PA32))),(PA33,((PA36,(Tarsius_syrichta,(Saimiri_sciureus,(Homo_sapiens,PA37)))),(PA35,(Lemur_catta,Notharctus_tenebrosus)))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,((PA38,((PA43,(((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(Leptictis_dakotensis,(Erinaceus_europaeus,(PA24,((Solenodon_paradoxus,PA23),(Sorex_araneus,(Talpa_europaea,PA25))))))),(Rhynchocyon_cirnei,PA44))),((PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(Orycteropus_afer,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA5,((PA6,(PA7,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),((Archaeotherium_mortoni,Mesonyx_obtusidens),(Rodhocetus_balochistanensis,((PA10,((Sus_scrofa,(PA12,(PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15)))))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,((PA22,(Canis_lupus,Felis_silvestris)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))))))),((PA26,(PA27,(PA28,(((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(PA29,(Rattus_norvegicus,PA30))),((Tribosphenomys_minutus,(Cocomys_lingchaensis,Paramys_delicatus)),(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),(Ptilocercus_lowii,(Tupaia_glis,PA32))),(PA33,(PA35,(((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36)),(Lemur_catta,Notharctus_tenebrosus)))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));

[“craniodental only” PAs corrected for the O'Leary et al. character ontology: strict consensus of 12 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000,PA3:0.000000,PA2:0.000000,PA1:0.000000,(PA5:0.000000,(PA38:0.000000,(PA40:0.000000,(PA43:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000,((Amblysomus_hottentotus:0.000000,Echinops_telfairi:0.000000,PA45:0.000000)0.336000:0.000000,(Leptictis_dakotensis:0.000000,(Erinaceus_europaeus:0.000000,(PA24:0.000000,(Solenodon_paradoxus:0.000000,PA23:0.000000)0.028000:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000,PA25:0.000000)0.744000:0.000000)0.032000:0.000000)0.120000:0.000000)0.012000:0.000000)0.000000:0.000000)0.004000:0.000000)0.000000:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.164000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.144000:0.000000)0.076000:0.000000)0.032000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.176000:0.000000)0.012000:0.000000)0.132000:0.000000,(Orycteropus_afer:0.000000,(((Dasypus_novemcinctus:0.000000,PA39:0.000000)0.116000:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.996000:0.000000)0.472000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.964000:0.000000)0.300000:0.000000)0.012000:0.000000)0.000000:0.000000)0.000000:0.000000,(PA6:0.000000,(PA7:0.000000,(PA8:0.000000,((((Apheliscus_insidiosus:0.000000,Protungulatum_donnae:0.000000)0.028000:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.280000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.248000:0.000000)0.060000:0.000000)0.180000:0.000000,((Archaeotherium_mortoni:0.000000,Mesonyx_obtusidens:0.000000)0.100000:0.000000,(Rodhocetus_balochistanensis:0.000000,((PA10:0.000000,(PA11:0.000000,(Sus_scrofa:0.000000,PA12:0.000000,(PA13:0.000000,(Hippopotamus_amphibius:0.000000,PA14:0.000000,((Artiocetus_clavis:0.000000,Basilosaurus_cetoides:0.000000)0.236000:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000,PA15:0.000000)0.980000:0.000000)0.388000:0.000000)0.196000:0.000000)0.004000:0.000000)0.004000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.036000:0.000000)0.020000:0.000000)0.000000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.440000:0.000000)0.056000:0.000000)0.008000:0.000000)0.476000:0.000000)0.004000:0.000000,(PA9:0.000000,(((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,(((Myotis_lucifugus:0.000000,PA17:0.000000)0.228000:0.000000,(Pteronotus_parnellii:0.000000,PA18:0.000000)0.188000:0.000000)0.320000:0.000000,((Nycteris_thebaica:0.000000,PA19:0.000000)0.148000:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.552000:0.000000)0.792000:0.000000)0.708000:0.000000)0.344000:0.000000,(PA16:0.000000,(Pteropus_giganteus:0.000000,PA20:0.000000)0.284000:0.000000)0.160000:0.000000)0.992000:0.000000)0.408000:0.000000)0.000000:0.000000,(PA21:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000,PA22:0.000000)0.936000:0.000000,(Sinopa_rapax:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.692000:0.000000)0.900000:0.000000)0.120000:0.000000)0.000000:0.000000)0.000000:0.000000,(PA26:0.000000,(PA27:0.000000,(PA28:0.000000,((PA29:0.000000,(Ictidomys_tridecemlineatus:0.000000,PA31:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.476000:0.000000)0.136000:0.000000,(Rattus_norvegicus:0.000000,PA30:0.000000)0.240000:0.000000)0.172000:0.000000,(Tribosphenomys_minutus:0.000000,(Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.688000:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.776000:0.000000)1.000000:0.000000)0.412000:0.000000)0.316000:0.000000)0.380000:0.000000)1.000000:0.000000)0.036000:0.000000,(PA33:0.000000,((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000,PA34:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,(Tupaia_glis:0.000000,PA32:0.000000)0.024000:0.000000)1.000000:0.000000)0.004000:0.000000,(PA35:0.000000,(Tarsius_syrichta:0.000000,PA36:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000,PA37:0.000000)1.000000:0.000000)0.468000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000)0.292000:0.000000)1.000000:0.000000)0.036000:0.000000)0.504000:0.000000)0.000000:0.000000)0.212000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.988000:0.000000)0.992000:0.000000,((Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000)0.376000:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.484000:0.000000)0.472000:0.000000)0.412000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.952000:0.000000)1.000000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“craniodental only” PAs corrected for the O'Leary et al. character ontology: strict consensus of 12 MPTs - nodal values = "transfer bootstrap expectation" values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000,PA3:0.000000,PA2:0.000000,PA1:0.000000,(PA5:0.000000,(PA38:0.000000,(PA40:0.000000,(PA43:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000,((Amblysomus_hottentotus:0.000000,Echinops_telfairi:0.000000,PA45:0.000000)0.370000:0.000000,(Leptictis_dakotensis:0.000000,(Erinaceus_europaeus:0.000000,(PA24:0.000000,(Solenodon_paradoxus:0.000000,PA23:0.000000)0.028000:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000,PA25:0.000000)0.750000:0.000000)0.399200:0.000000)0.395333:0.000000)0.342857:0.000000)0.251200:0.000000)0.215000:0.000000)0.198462:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.164000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.144000:0.000000)0.174000:0.000000)0.187000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.176000:0.000000)0.246667:0.000000)0.268571:0.000000,(Orycteropus_afer:0.000000,(((Dasypus_novemcinctus:0.000000,PA39:0.000000)0.116000:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.996000:0.000000)0.648000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.964000:0.000000)0.588800:0.000000)0.492667:0.000000)0.155724:0.000000)0.150800:0.000000,(PA6:0.000000,(PA7:0.000000,(PA8:0.000000,((((Apheliscus_insidiosus:0.000000,Protungulatum_donnae:0.000000)0.028000:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.280000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.248000:0.000000)0.285333:0.000000)0.359200:0.000000,((Archaeotherium_mortoni:0.000000,Mesonyx_obtusidens:0.000000)0.100000:0.000000,(Rodhocetus_balochistanensis:0.000000,((PA10:0.000000,(PA11:0.000000,(Sus_scrofa:0.000000,PA12:0.000000,(PA13:0.000000,(Hippopotamus_amphibius:0.000000,PA14:0.000000,((Artiocetus_clavis:0.000000,Basilosaurus_cetoides:0.000000)0.236000:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000,PA15:0.000000)0.980000:0.000000)0.728000:0.000000)0.576000:0.000000)0.494286:0.000000)0.386222:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.036000:0.000000)0.419333:0.000000)0.462462:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.440000:0.000000)0.537600:0.000000)0.560250:0.000000)0.602889:0.000000)0.469000:0.000000,(PA9:0.000000,(((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,(((Myotis_lucifugus:0.000000,PA17:0.000000)0.228000:0.000000,(Pteronotus_parnellii:0.000000,PA18:0.000000)0.188000:0.000000)0.364000:0.000000,((Nycteris_thebaica:0.000000,PA19:0.000000)0.148000:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.552000:0.000000)0.816000:0.000000)0.824000:0.000000)0.826222:0.000000,(PA16:0.000000,(Pteropus_giganteus:0.000000,PA20:0.000000)0.284000:0.000000)0.240000:0.000000)0.998000:0.000000)0.952615:0.000000)0.404526:0.000000,(PA21:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000,PA22:0.000000)0.936000:0.000000,(Sinopa_rapax:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.846000:0.000000)0.946400:0.000000)0.811333:0.000000)0.339391:0.000000)0.332255:0.000000,(PA26:0.000000,(PA27:0.000000,(PA28:0.000000,((PA29:0.000000,(Ictidomys_tridecemlineatus:0.000000,PA31:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.476000:0.000000)0.273333:0.000000,(Rattus_norvegicus:0.000000,PA30:0.000000)0.240000:0.000000)0.334667:0.000000,(Tribosphenomys_minutus:0.000000,(Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.688000:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.776000:0.000000)1.000000:0.000000)0.804000:0.000000)0.662000:0.000000)0.952308:0.000000)1.000000:0.000000)0.935733:0.000000,(PA33:0.000000,((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000,PA34:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,(Tupaia_glis:0.000000,PA32:0.000000)0.024000:0.000000)1.000000:0.000000)0.402400:0.000000,(PA35:0.000000,(Tarsius_syrichta:0.000000,PA36:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000,PA37:0.000000)1.000000:0.000000)0.783000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000)0.292000:0.000000)1.000000:0.000000)0.693143:0.000000)0.739613:0.000000)0.172735:0.000000)0.443765:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.988000:0.000000)0.992000:0.000000,((Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000)0.376000:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.484000:0.000000)0.601333:0.000000)0.707000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.952000:0.000000)1.000000:0.000000)1.000000:0.000000));

[“craniodental only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: 8 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(((PA7,(PA8,((PA21,((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))),(PA6,((PA26,(PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))))),((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))),(PA5,(PA38,(((PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(PA43,(Orycteropus_afer,(Rhynchocyon_cirnei,PA44)))),(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(((PA7,(PA8,((PA21,((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))),(PA6,((PA26,(PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))))),((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))),(PA5,((PA38,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA40,(PA43,(Orycteropus_afer,(Rhynchocyon_cirnei,PA44)))))),(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((((Solenodon_paradoxus,(Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea))),(Echinops_telfairi,Erinaceus_europaeus)),((PA6,((PA7,((PA21,((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))))),(PA8,((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))),(PA26,((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA33,(((Cynocephalus_volans,Galeopterus_variegatus),(Ptilocercus_lowii,Tupaia_glis)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus)))))))),(PA5,(PA38,(((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA40,((Orycteropus_afer,PA43),(Rhynchocyon_cirnei,PA44)))),(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(((PA7,(PA8,((PA21,((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))),(PA6,((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA26,((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))))),(PA5,(PA38,((PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),((Orycteropus_afer,PA43),(Rhynchocyon_cirnei,PA44)))),(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((((Solenodon_paradoxus,(Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea))),(Echinops_telfairi,Erinaceus_europaeus)),((PA6,((PA7,((PA8,(PA21,((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus)))))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))))),(PA26,((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA33,(((Cynocephalus_volans,Galeopterus_variegatus),(Ptilocercus_lowii,Tupaia_glis)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus)))))))),(PA5,(PA38,(((PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(Orycteropus_afer,(PA43,(Rhynchocyon_cirnei,PA44)))),(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((((Solenodon_paradoxus,(Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea))),(Echinops_telfairi,Erinaceus_europaeus)),((PA6,((PA7,((PA21,((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(PA8,((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))),(PA26,((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA33,(((Cynocephalus_volans,Galeopterus_variegatus),(Ptilocercus_lowii,Tupaia_glis)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus)))))))),(PA5,((PA38,((PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(Orycteropus_afer,(PA43,(Rhynchocyon_cirnei,PA44))))),(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(((PA7,((PA8,(PA21,((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))))),(PA6,((PA26,(PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))))),((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))),(PA5,((PA38,(PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(Orycteropus_afer,(PA43,(Rhynchocyon_cirnei,PA44)))))),(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((((Solenodon_paradoxus,(Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea))),(Echinops_telfairi,Erinaceus_europaeus)),(((PA7,((PA21,((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))))),(PA8,((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))),(PA6,((PA26,(PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))))),(PA33,(((Cynocephalus_volans,Galeopterus_variegatus),(Ptilocercus_lowii,Tupaia_glis)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus))))))),(PA5,((PA38,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA40,((Orycteropus_afer,PA43),(Rhynchocyon_cirnei,PA44))))),(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));

[“craniodental only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: strict consensus of 8 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.968000:0.000000,((Eomaia_scansoria:0.000000,(((((Solenodon_paradoxus:0.000000,(Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.500000:0.000000)0.784000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.736000:0.000000)0.352000:0.000000,((PA6:0.000000,(PA7:0.000000,(PA8:0.000000,(PA21:0.000000,((Leptictis_dakotensis:0.000000,(Protungulatum_donnae:0.000000,(Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.344000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.260000:0.000000)0.092000:0.000000)0.140000:0.000000)0.104000:0.000000)0.008000:0.000000,(Sinopa_rapax:0.000000,(Canis_lupus:0.000000,Felis_silvestris:0.000000)0.976000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.968000:0.000000)0.012000:0.000000)0.004000:0.000000,((PA9:0.000000,(PA10:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.992000:0.000000)0.104000:0.000000)0.720000:0.000000)0.088000:0.000000)0.020000:0.000000)0.016000:0.000000)0.036000:0.000000)0.024000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.924000:0.000000)0.020000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.580000:0.000000)0.004000:0.000000)0.460000:0.000000)0.000000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.664000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.708000:0.000000)0.984000:0.000000)1.000000:0.000000)0.992000:0.000000)1.000000:0.000000)0.004000:0.000000)0.004000:0.000000)0.004000:0.000000,(PA26:0.000000,(PA27:0.000000,(PA28:0.000000,((Ictidomys_tridecemlineatus:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.868000:0.000000)0.340000:0.000000,(Rattus_norvegicus:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.368000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.688000:0.000000)1.000000:0.000000)0.324000:0.000000)0.288000:0.000000)0.100000:0.000000)0.084000:0.000000)0.564000:0.000000)1.000000:0.000000)0.128000:0.000000,(PA33:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.992000:0.000000)0.428000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.148000:0.000000)0.332000:0.000000)0.004000:0.000000,(PA5:0.000000,(PA38:0.000000,(PA40:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.124000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.084000:0.000000)0.088000:0.000000)0.016000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.196000:0.000000)0.004000:0.000000)0.056000:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000)0.912000:0.000000)0.156000:0.000000)0.004000:0.000000,(((Dasypus_novemcinctus:0.000000,PA39:0.000000)0.136000:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)1.000000:0.000000)0.548000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.964000:0.000000)0.304000:0.000000)0.008000:0.000000)0.000000:0.000000)0.000000:0.000000)0.120000:0.000000,(PA3:0.000000,PA1:0.000000)0.044000:0.000000)0.228000:0.000000,(Dromiciops_gliroides:0.000000,(PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.972000:0.000000)0.000000:0.000000)0.996000:0.000000)0.364000:0.000000,(Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.724000:0.000000)0.284000:0.000000)0.288000:0.000000)0.960000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.820000:0.000000)0.896000:0.000000)1.000000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“craniodental only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: strict consensus of 8 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.968000:0.000000,((Eomaia_scansoria:0.000000,(((((Solenodon_paradoxus:0.000000,(Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.500000:0.000000)0.852000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.736000:0.000000)0.651200:0.000000,((PA6:0.000000,(PA7:0.000000,(PA8:0.000000,(PA21:0.000000,((Leptictis_dakotensis:0.000000,(Protungulatum_donnae:0.000000,(Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.344000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.260000:0.000000)0.317333:0.000000)0.348000:0.000000)0.318400:0.000000)0.266667:0.000000,(Sinopa_rapax:0.000000,(Canis_lupus:0.000000,Felis_silvestris:0.000000)0.976000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.976000:0.000000)0.376364:0.000000)0.345333:0.000000,((PA9:0.000000,(PA10:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.992000:0.000000)0.550000:0.000000)0.837333:0.000000)0.651000:0.000000)0.526400:0.000000)0.449333:0.000000)0.400000:0.000000)0.361500:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.924000:0.000000)0.456400:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.580000:0.000000)0.550667:0.000000)0.596000:0.000000)0.553429:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.664000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.708000:0.000000)0.988000:0.000000)1.000000:0.000000)0.998667:0.000000)1.000000:0.000000)0.458182:0.000000)0.286333:0.000000)0.278703:0.000000,(PA26:0.000000,(PA27:0.000000,(PA28:0.000000,((Ictidomys_tridecemlineatus:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.868000:0.000000)0.642000:0.000000,(Rattus_norvegicus:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.368000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.688000:0.000000)1.000000:0.000000)0.774667:0.000000)0.706000:0.000000)0.526000:0.000000)0.572571:0.000000)0.956400:0.000000)1.000000:0.000000)0.927333:0.000000,(PA33:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.992000:0.000000)0.710000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.597333:0.000000)0.651478:0.000000)0.216000:0.000000,(PA5:0.000000,(PA38:0.000000,(PA40:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.124000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.084000:0.000000)0.142000:0.000000)0.142000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.196000:0.000000)0.164667:0.000000)0.171429:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000)0.912000:0.000000)0.414667:0.000000)0.169333:0.000000,(((Dasypus_novemcinctus:0.000000,PA39:0.000000)0.136000:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)1.000000:0.000000)0.698667:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.964000:0.000000)0.627200:0.000000)0.197263:0.000000)0.187400:0.000000)0.412000:0.000000)0.565500:0.000000,(PA3:0.000000,PA1:0.000000)0.044000:0.000000)0.621714:0.000000,(Dromiciops_gliroides:0.000000,(PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.972000:0.000000)0.500000:0.000000)0.997333:0.000000)0.750800:0.000000,(Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.724000:0.000000)0.552000:0.000000)0.877714:0.000000)0.976000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.820000:0.000000)0.948000:0.000000)1.000000:0.000000)1.000000:0.000000)-1.#IND00:0.000000);

[“craniodental only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: 2 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((PA1,(PA3,(PA5,(PA6,((PA7,(((((Solenodon_paradoxus,(Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea))),(Echinops_telfairi,Erinaceus_europaeus)),(Leptictis_dakotensis,Rhynchocyon_cirnei)),((((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)),(PA38,(Orycteropus_afer,PA40)))),(((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((PA1,(PA3,(PA5,(PA6,((PA7,(((((Solenodon_paradoxus,(Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea))),(Echinops_telfairi,Erinaceus_europaeus)),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(((PA38,(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla)))),(Orycteropus_afer,PA40)),(Manis_pentadactyla,Metacheiromys_marshi))),(((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));

[“craniodental only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: strict consensus of 2 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.988000:0.000000,((Eomaia_scansoria:0.000000,(((PA3:0.000000,PA1:0.000000,(PA5:0.000000,(PA6:0.000000,((PA7:0.000000,(((((Solenodon_paradoxus:0.000000,(Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.496000:0.000000)0.804000:0.000000)0.048000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.840000:0.000000)0.324000:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.336000:0.000000)0.044000:0.000000,(PA38:0.000000,(Dasypus_novemcinctus:0.000000,PA39:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.996000:0.000000)0.500000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.968000:0.000000,(Orycteropus_afer:0.000000,PA40:0.000000)0.156000:0.000000)0.332000:0.000000)0.004000:0.000000,(((Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.256000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.392000:0.000000)0.076000:0.000000)0.292000:0.000000,(Protungulatum_donnae:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)0.996000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.728000:0.000000)1.000000:0.000000)0.048000:0.000000)0.024000:0.000000,(((Procavia_capensis:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.992000:0.000000)0.100000:0.000000)0.604000:0.000000)0.076000:0.000000)0.016000:0.000000)0.052000:0.000000)0.024000:0.000000)0.012000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.980000:0.000000)0.020000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.604000:0.000000)0.260000:0.000000)0.008000:0.000000,((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.028000:0.000000,(Moeritherium_lyonsi:0.000000,(Loxodonta_africana:0.000000,Trichechus_manatus:0.000000)0.180000:0.000000)0.320000:0.000000)0.036000:0.000000)0.164000:0.000000,((Rattus_norvegicus:0.000000,((Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.836000:0.000000,(Cocomys_lingchaensis:0.000000,Ictidomys_tridecemlineatus:0.000000)0.368000:0.000000)0.204000:0.000000)0.116000:0.000000,(Paramys_delicatus:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.852000:0.000000)1.000000:0.000000)0.424000:0.000000)0.296000:0.000000)0.056000:0.000000)1.000000:0.000000)0.092000:0.000000)0.128000:0.000000)0.000000:0.000000)0.000000:0.000000,(PA26:0.000000,(((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.664000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.752000:0.000000)0.972000:0.000000)1.000000:0.000000)0.992000:0.000000)1.000000:0.000000)0.040000:0.000000,((Notharctus_tenebrosus:0.000000,(Lemur_catta:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.980000:0.000000)0.468000:0.000000)0.132000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.116000:0.000000)0.020000:0.000000)0.052000:0.000000)0.000000:0.000000)0.000000:0.000000)0.084000:0.000000)0.220000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.960000:0.000000)0.976000:0.000000)0.504000:0.000000,(Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.732000:0.000000)0.256000:0.000000)0.236000:0.000000)0.976000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.856000:0.000000)0.888000:0.000000)1.000000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“craniodental only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: strict consensus of 2 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.988000:0.000000,((Eomaia_scansoria:0.000000,(((PA3:0.000000,PA1:0.000000,(PA5:0.000000,(PA6:0.000000,((PA7:0.000000,(((((Solenodon_paradoxus:0.000000,(Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.496000:0.000000)0.870000:0.000000)0.605333:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.840000:0.000000)0.610400:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.336000:0.000000)0.457714:0.000000,(PA38:0.000000,(Dasypus_novemcinctus:0.000000,PA39:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.996000:0.000000)0.666667:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.968000:0.000000,(Orycteropus_afer:0.000000,PA40:0.000000)0.156000:0.000000)0.596500:0.000000)0.332750:0.000000,(((Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.256000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.392000:0.000000)0.361333:0.000000)0.439000:0.000000,(Protungulatum_donnae:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)0.996000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.909333:0.000000)1.000000:0.000000)0.809600:0.000000)0.424800:0.000000,(((Procavia_capensis:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.992000:0.000000)0.548000:0.000000)0.748000:0.000000)0.581000:0.000000)0.476800:0.000000)0.418000:0.000000)0.366857:0.000000)0.350500:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.980000:0.000000)0.395200:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.604000:0.000000)0.463333:0.000000)0.444923:0.000000,((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.028000:0.000000,(Moeritherium_lyonsi:0.000000,(Loxodonta_africana:0.000000,Trichechus_manatus:0.000000)0.180000:0.000000)0.514000:0.000000)0.479000:0.000000)0.425778:0.000000,((Rattus_norvegicus:0.000000,((Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.836000:0.000000,(Cocomys_lingchaensis:0.000000,Ictidomys_tridecemlineatus:0.000000)0.368000:0.000000)0.521333:0.000000)0.511000:0.000000,(Paramys_delicatus:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.852000:0.000000)1.000000:0.000000)0.808000:0.000000)0.737000:0.000000)0.604000:0.000000)1.000000:0.000000)0.486345:0.000000)0.426500:0.000000)0.332432:0.000000)0.332333:0.000000,(PA26:0.000000,(((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.664000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.752000:0.000000)0.980000:0.000000)1.000000:0.000000)0.998667:0.000000)1.000000:0.000000)0.786667:0.000000,((Notharctus_tenebrosus:0.000000,(Lemur_catta:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.980000:0.000000)0.724000:0.000000)0.710667:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.705333:0.000000)0.485000:0.000000)0.516706:0.000000)0.516222:0.000000)0.546588:0.000000)0.580750:0.000000)0.646571:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.960000:0.000000)0.984000:0.000000)0.804400:0.000000,(Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.732000:0.000000)0.542000:0.000000)0.878286:0.000000)0.985333:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.856000:0.000000)0.944000:0.000000)1.000000:0.000000)1.000000:0.000000)-1.#IND00:0.000000);

[“dental only" PAs corrected for the O'Leary et al. character ontology: 9 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((PA3,((PA5,(PA6,((PA7,((PA23,((PA24,((Amblysomus_hottentotus,Talpa_europaea),(Sorex_araneus,PA25))),(Solenodon_paradoxus,(Erinaceus_europaeus,(Echinops_telfairi,PA45))))),(PA8,(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,(PA17,(Pteronotus_parnellii,PA18))),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))))),(((Leptictis_dakotensis,((Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))),(Sinopa_rapax,((Canis_lupus,(PA22,(Felis_silvestris,PA21))),(Vulpavus_ovatus,Vulpavus_profectus))))),((PA39,(PA38,((PA41,(PA42,(Procavia_capensis,((Mesohippus_bairdi,(Equus_caballus,(PA10,((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),((PA12,(PA15,(PA13,(Bos_taurus,PA14)))),(Lama_glama,PA11)))))),(Thomashuxleya_externa_comb,(Carodnia_vieirai,(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus)))))))),(PA40,(PA43,(Rhynchocyon_cirnei,PA44)))))),(Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))),((PA27,(PA28,((Rattus_norvegicus,(Ictidomys_tridecemlineatus,(PA30,(PA31,(PA29,(Castor_canadensis,Cavia_porcellus)))))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(PA26,((PA33,(Galeopterus_variegatus,(Cynocephalus_volans,PA34))),((Notharctus_tenebrosus,(((PA37,(Homo_sapiens,Saimiri_sciureus)),(Tarsius_syrichta,PA36)),(Lemur_catta,PA35))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))),(PA2,PA1))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((PA3,((PA5,(PA6,((PA7,((PA23,((PA24,((Amblysomus_hottentotus,Talpa_europaea),(Sorex_araneus,PA25))),(Solenodon_paradoxus,(Erinaceus_europaeus,(Echinops_telfairi,PA45))))),(PA8,(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,(PA17,(Pteronotus_parnellii,PA18))),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))))),(((Leptictis_dakotensis,((Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))),(Sinopa_rapax,((Canis_lupus,(PA21,(Felis_silvestris,PA22))),(Vulpavus_ovatus,Vulpavus_profectus))))),((PA39,(PA38,((PA41,(PA42,(Procavia_capensis,((Mesohippus_bairdi,(Equus_caballus,(PA10,((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),((Bos_taurus,(PA15,(PA14,PA13))),(PA12,(Lama_glama,PA11))))))),(Thomashuxleya_externa_comb,(Carodnia_vieirai,(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus)))))))),(PA40,(PA43,(Rhynchocyon_cirnei,PA44)))))),(Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))),((PA27,(PA28,((Rattus_norvegicus,(Ictidomys_tridecemlineatus,(PA31,((Castor_canadensis,Cavia_porcellus),(PA30,PA29))))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(PA26,((PA33,(Galeopterus_variegatus,(Cynocephalus_volans,PA34))),((Notharctus_tenebrosus,((PA36,(Tarsius_syrichta,(Homo_sapiens,(Saimiri_sciureus,PA37)))),(Lemur_catta,PA35))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))),(PA2,PA1))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((PA5,((PA6,((PA7,((PA24,(Sorex_araneus,(PA25,(Amblysomus_hottentotus,Talpa_europaea)))),(PA23,(Solenodon_paradoxus,(Erinaceus_europaeus,(Echinops_telfairi,PA45)))))),(PA8,(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))))),(((Leptictis_dakotensis,((Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))),(Sinopa_rapax,((Canis_lupus,(Felis_silvestris,(PA22,PA21))),(Vulpavus_ovatus,Vulpavus_profectus))))),((PA39,(PA38,((PA41,(PA42,(Procavia_capensis,((Mesohippus_bairdi,(Equus_caballus,(PA10,((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(PA11,(Lama_glama,(PA12,(PA14,(PA13,(Bos_taurus,PA15)))))))))),(Thomashuxleya_externa_comb,(Carodnia_vieirai,(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus)))))))),(PA40,(PA43,(Rhynchocyon_cirnei,PA44)))))),(Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))),((PA27,(PA28,((Rattus_norvegicus,(Ictidomys_tridecemlineatus,((Castor_canadensis,Cavia_porcellus),(PA30,(PA31,PA29))))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(PA26,((PA33,(Cynocephalus_volans,(Galeopterus_variegatus,PA34))),((Notharctus_tenebrosus,((PA36,(Tarsius_syrichta,(Saimiri_sciureus,(Homo_sapiens,PA37)))),(Lemur_catta,PA35))),(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))),(PA3,(PA2,PA1))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((PA3,(PA1,(PA2,(PA5,((PA7,(((PA24,(PA25,(Sorex_araneus,(Amblysomus_hottentotus,Talpa_europaea)))),(PA23,(Solenodon_paradoxus,(Erinaceus_europaeus,(Echinops_telfairi,PA45))))),(PA8,(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))))),(((Leptictis_dakotensis,((Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))),(Sinopa_rapax,((Canis_lupus,(PA21,(Felis_silvestris,PA22))),(Vulpavus_ovatus,Vulpavus_profectus))))),((((PA40,(PA41,(PA42,(Procavia_capensis,((Mesohippus_bairdi,(Equus_caballus,(PA10,((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Lama_glama,(PA11,(PA12,(Bos_taurus,(PA13,(PA15,PA14)))))))))),(Thomashuxleya_externa_comb,(Carodnia_vieirai,(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))))))))),(PA43,(Rhynchocyon_cirnei,PA44))),(PA39,PA38)),(Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA6,((PA26,(PA27,(PA28,((Rattus_norvegicus,(Ictidomys_tridecemlineatus,((PA30,(Castor_canadensis,Cavia_porcellus)),(PA31,PA29)))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),((PA33,(PA34,(Cynocephalus_volans,Galeopterus_variegatus))),((Notharctus_tenebrosus,((PA36,(Tarsius_syrichta,(Homo_sapiens,(Saimiri_sciureus,PA37)))),(Lemur_catta,PA35))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((PA3,(PA2,(PA1,(PA5,((PA7,((PA23,((PA24,(PA25,(Sorex_araneus,(Amblysomus_hottentotus,Talpa_europaea)))),(Solenodon_paradoxus,(Erinaceus_europaeus,(Echinops_telfairi,PA45))))),(PA8,(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),((PA17,(Myotis_lucifugus,(Pteronotus_parnellii,PA18))),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))))),((PA6,((Leptictis_dakotensis,((Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))),(Sinopa_rapax,((Canis_lupus,(PA21,(Felis_silvestris,PA22))),(Vulpavus_ovatus,Vulpavus_profectus))))),((PA39,(PA38,(PA40,((PA41,(PA42,(Procavia_capensis,((Mesohippus_bairdi,(Equus_caballus,(PA10,((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),((Bos_taurus,(PA13,(PA15,PA14))),(PA12,(Lama_glama,PA11))))))),(Thomashuxleya_externa_comb,(Carodnia_vieirai,(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus)))))))),(PA43,(Rhynchocyon_cirnei,PA44)))))),(Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))),((PA27,(PA28,((Rattus_norvegicus,(Ictidomys_tridecemlineatus,((PA30,(Castor_canadensis,Cavia_porcellus)),(PA31,PA29)))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(PA26,((PA33,(Galeopterus_variegatus,(Cynocephalus_volans,PA34))),((Notharctus_tenebrosus,((PA36,(Tarsius_syrichta,(PA37,(Homo_sapiens,Saimiri_sciureus)))),(Lemur_catta,PA35))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((PA2,((PA5,((((PA23,(PA24,(PA25,(Sorex_araneus,(Amblysomus_hottentotus,Talpa_europaea))))),(Solenodon_paradoxus,(Erinaceus_europaeus,(Echinops_telfairi,PA45)))),(PA7,(PA8,(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),((PA17,(Myotis_lucifugus,(Pteronotus_parnellii,PA18))),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))))),(((Leptictis_dakotensis,((Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))),(Sinopa_rapax,((Canis_lupus,(PA21,(Felis_silvestris,PA22))),(Vulpavus_ovatus,Vulpavus_profectus))))),(((PA40,((PA41,(PA42,(Procavia_capensis,((Mesohippus_bairdi,(Equus_caballus,(PA10,((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),((PA12,(PA14,(Bos_taurus,(PA15,PA13)))),(Lama_glama,PA11)))))),(Thomashuxleya_externa_comb,(Carodnia_vieirai,(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus)))))))),(PA43,(Rhynchocyon_cirnei,PA44)))),(PA39,PA38)),(Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA6,((PA27,(PA28,((Rattus_norvegicus,(Ictidomys_tridecemlineatus,((Castor_canadensis,Cavia_porcellus),(PA29,(PA30,PA31))))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(PA26,((PA33,(Galeopterus_variegatus,(Cynocephalus_volans,PA34))),((Notharctus_tenebrosus,((PA36,(Tarsius_syrichta,(Homo_sapiens,(Saimiri_sciureus,PA37)))),(Lemur_catta,PA35))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))),(PA3,PA1))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((PA5,((PA7,(((PA23,(PA24,((Amblysomus_hottentotus,Talpa_europaea),(Sorex_araneus,PA25)))),(Solenodon_paradoxus,(Erinaceus_europaeus,(Echinops_telfairi,PA45)))),(PA8,(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,(PA17,(Pteronotus_parnellii,PA18))),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))))),(PA6,(((Leptictis_dakotensis,((Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))),(Sinopa_rapax,((Canis_lupus,(Felis_silvestris,(PA22,PA21))),(Vulpavus_ovatus,Vulpavus_profectus))))),((PA38,(PA39,((PA40,(PA41,(PA42,(Procavia_capensis,((Mesohippus_bairdi,(Equus_caballus,(PA10,((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Lama_glama,(PA11,(PA12,(PA13,(PA14,(Bos_taurus,PA15)))))))))),(Thomashuxleya_externa_comb,(Carodnia_vieirai,(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))))))))),(PA43,(Rhynchocyon_cirnei,PA44))))),(Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA26,((PA27,(PA28,((Rattus_norvegicus,(Ictidomys_tridecemlineatus,((Castor_canadensis,Cavia_porcellus),(PA31,(PA30,PA29))))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),((PA33,(Galeopterus_variegatus,(Cynocephalus_volans,PA34))),((Notharctus_tenebrosus,((Tarsius_syrichta,(PA36,(PA37,(Homo_sapiens,Saimiri_sciureus)))),(Lemur_catta,PA35))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))),(PA1,(PA3,PA2))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((PA2,((PA5,((PA7,(((PA23,(PA24,((Amblysomus_hottentotus,Talpa_europaea),(Sorex_araneus,PA25)))),(Solenodon_paradoxus,(Erinaceus_europaeus,(Echinops_telfairi,PA45)))),(PA8,(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,(PA17,(Pteronotus_parnellii,PA18))),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))))),(PA6,(((Leptictis_dakotensis,((Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))),(Sinopa_rapax,((Canis_lupus,(Felis_silvestris,(PA22,PA21))),(Vulpavus_ovatus,Vulpavus_profectus))))),((PA38,(PA39,((PA40,(PA41,(PA42,(Procavia_capensis,((Mesohippus_bairdi,(Equus_caballus,(PA10,((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),((PA13,(PA15,(Bos_taurus,PA14))),(PA12,(Lama_glama,PA11))))))),(Thomashuxleya_externa_comb,(Carodnia_vieirai,(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))))))))),(PA43,(Rhynchocyon_cirnei,PA44))))),(Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA26,((PA27,(PA28,((Rattus_norvegicus,(Ictidomys_tridecemlineatus,((Castor_canadensis,Cavia_porcellus),(PA31,(PA30,PA29))))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),((PA33,(Galeopterus_variegatus,(Cynocephalus_volans,PA34))),((Notharctus_tenebrosus,((Tarsius_syrichta,(PA36,(PA37,(Homo_sapiens,Saimiri_sciureus)))),(Lemur_catta,PA35))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))),(PA3,PA1))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((PA5,((((PA24,(Sorex_araneus,(PA25,(Amblysomus_hottentotus,Talpa_europaea)))),(PA23,(Solenodon_paradoxus,(Erinaceus_europaeus,(Echinops_telfairi,PA45))))),(PA7,(PA8,(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),((PA17,(Myotis_lucifugus,(Pteronotus_parnellii,PA18))),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))))),(PA6,(((Leptictis_dakotensis,((Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))),(Sinopa_rapax,((Canis_lupus,(PA22,(Felis_silvestris,PA21))),(Vulpavus_ovatus,Vulpavus_profectus))))),(((PA40,((PA41,(PA42,(Procavia_capensis,((Mesohippus_bairdi,(Equus_caballus,(PA10,((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),((PA12,(Bos_taurus,(PA15,(PA14,PA13)))),(Lama_glama,PA11)))))),(Thomashuxleya_externa_comb,(Carodnia_vieirai,(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus)))))))),(PA43,(Rhynchocyon_cirnei,PA44)))),(PA39,PA38)),(Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))),((PA26,(PA27,(PA28,((Rattus_norvegicus,(Ictidomys_tridecemlineatus,(PA29,((Castor_canadensis,Cavia_porcellus),(PA30,PA31))))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),((PA33,(Cynocephalus_volans,(Galeopterus_variegatus,PA34))),((Notharctus_tenebrosus,((Tarsius_syrichta,(PA36,(Saimiri_sciureus,(Homo_sapiens,PA37)))),(Lemur_catta,PA35))),(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))),(PA3,(PA2,PA1))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));

[“dental only” PAs corrected for the O'Leary et al. character ontology: strict consensus of 9 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.968000:0.000000,((Eomaia_scansoria:0.000000,(((PA3:0.000000,PA2:0.000000,PA1:0.000000,(PA5:0.000000,(PA6:0.000000,(PA7:0.000000,(PA23:0.000000,(PA24:0.000000,(Sorex_araneus:0.000000,PA25:0.000000,(Amblysomus_hottentotus:0.000000,Talpa_europaea:0.000000)0.184000:0.000000)0.180000:0.000000)0.104000:0.000000,(Solenodon_paradoxus:0.000000,(Erinaceus_europaeus:0.000000,(Echinops_telfairi:0.000000,PA45:0.000000)0.144000:0.000000)0.024000:0.000000)0.000000:0.000000)0.024000:0.000000,(PA8:0.000000,(PA9:0.000000,(((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,PA17:0.000000,(Pteronotus_parnellii:0.000000,PA18:0.000000)0.164000:0.000000)0.188000:0.000000,((Nycteris_thebaica:0.000000,PA19:0.000000)0.132000:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.564000:0.000000)0.696000:0.000000)0.612000:0.000000)0.212000:0.000000,(PA16:0.000000,(Pteropus_giganteus:0.000000,PA20:0.000000)0.104000:0.000000)0.112000:0.000000)0.992000:0.000000)0.596000:0.000000)0.040000:0.000000)0.000000:0.000000,((Leptictis_dakotensis:0.000000,((Protungulatum_donnae:0.000000,(Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.320000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.352000:0.000000)0.096000:0.000000)0.152000:0.000000)0.144000:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,(Felis_silvestris:0.000000,PA22:0.000000,PA21:0.000000)0.084000:0.000000)0.184000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.048000:0.000000)0.180000:0.000000)0.000000:0.000000)0.000000:0.000000,((PA39:0.000000,PA38:0.000000,(PA40:0.000000,(PA41:0.000000,(PA42:0.000000,(Procavia_capensis:0.000000,((Mesohippus_bairdi:0.000000,(Equus_caballus:0.000000,(PA10:0.000000,((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.952000:0.000000)0.096000:0.000000)0.536000:0.000000)0.060000:0.000000)0.012000:0.000000)0.036000:0.000000)0.004000:0.000000)0.000000:0.000000,(Lama_glama:0.000000,PA12:0.000000,PA11:0.000000,(Bos_taurus:0.000000,PA15:0.000000,PA14:0.000000,PA13:0.000000)0.192000:0.000000)0.100000:0.000000)0.000000:0.000000)0.000000:0.000000)0.000000:0.000000)0.196000:0.000000,(Thomashuxleya_externa_comb:0.000000,(Carodnia_vieirai:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.144000:0.000000)0.240000:0.000000)0.288000:0.000000)0.040000:0.000000)0.016000:0.000000)0.004000:0.000000)0.000000:0.000000)0.004000:0.000000,(PA43:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000)0.044000:0.000000)0.000000:0.000000)0.000000:0.000000)0.000000:0.000000,(Orycteropus_afer:0.000000,((Dasypus_novemcinctus:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.792000:0.000000)0.376000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.508000:0.000000)0.124000:0.000000)0.096000:0.000000)0.000000:0.000000)0.000000:0.000000,(PA26:0.000000,(PA27:0.000000,(PA28:0.000000,((Rattus_norvegicus:0.000000,(Ictidomys_tridecemlineatus:0.000000,(PA30:0.000000,PA31:0.000000,PA29:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.616000:0.000000)0.264000:0.000000)0.012000:0.000000)0.036000:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.312000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.800000:0.000000)1.000000:0.000000)0.432000:0.000000)0.324000:0.000000)0.036000:0.000000)0.032000:0.000000)0.964000:0.000000)0.012000:0.000000,((PA33:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000,PA34:0.000000)1.000000:0.000000)0.096000:0.000000,((Notharctus_tenebrosus:0.000000,((Tarsius_syrichta:0.000000,PA36:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000,PA37:0.000000)0.996000:0.000000)0.472000:0.000000,(Lemur_catta:0.000000,PA35:0.000000)0.240000:0.000000)0.200000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,(Tupaia_glis:0.000000,PA32:0.000000)0.032000:0.000000)1.000000:0.000000)0.060000:0.000000)0.032000:0.000000)0.028000:0.000000)0.004000:0.000000)0.040000:0.000000)0.516000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.936000:0.000000)0.992000:0.000000)0.388000:0.000000,(Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.796000:0.000000)0.316000:0.000000)0.360000:0.000000)0.968000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.828000:0.000000)0.944000:0.000000)1.000000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“dental only” PAs corrected for the O'Leary et al. character ontology: strict consensus of 9 MPTs - nodal values = "transfer bootstrap expectation" values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.968000:0.000000,((Eomaia_scansoria:0.000000,(((PA3:0.000000,PA2:0.000000,PA1:0.000000,(PA5:0.000000,(PA6:0.000000,(PA7:0.000000,(PA23:0.000000,(PA24:0.000000,(Sorex_araneus:0.000000,PA25:0.000000,(Amblysomus_hottentotus:0.000000,Talpa_europaea:0.000000)0.184000:0.000000)0.425333:0.000000)0.369000:0.000000,(Solenodon_paradoxus:0.000000,(Erinaceus_europaeus:0.000000,(Echinops_telfairi:0.000000,PA45:0.000000)0.144000:0.000000)0.098000:0.000000)0.065333:0.000000)0.236000:0.000000,(PA8:0.000000,(PA9:0.000000,(((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,PA17:0.000000,(Pteronotus_parnellii:0.000000,PA18:0.000000)0.164000:0.000000)0.232000:0.000000,((Nycteris_thebaica:0.000000,PA19:0.000000)0.132000:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.564000:0.000000)0.726667:0.000000)0.786857:0.000000)0.792000:0.000000,(PA16:0.000000,(Pteropus_giganteus:0.000000,PA20:0.000000)0.104000:0.000000)0.138000:0.000000)0.996000:0.000000)0.965538:0.000000)0.899429:0.000000)0.504000:0.000000,((Leptictis_dakotensis:0.000000,((Protungulatum_donnae:0.000000,(Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.320000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.352000:0.000000)0.326667:0.000000)0.377000:0.000000)0.356800:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,(Felis_silvestris:0.000000,PA22:0.000000,PA21:0.000000)0.136000:0.000000)0.194667:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.324800:0.000000)0.346667:0.000000)0.244000:0.000000)0.225231:0.000000,((PA39:0.000000,PA38:0.000000,(PA40:0.000000,(PA41:0.000000,(PA42:0.000000,(Procavia_capensis:0.000000,((Mesohippus_bairdi:0.000000,(Equus_caballus:0.000000,(PA10:0.000000,((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.952000:0.000000)0.526000:0.000000)0.694667:0.000000)0.537000:0.000000)0.439200:0.000000)0.380667:0.000000)0.327429:0.000000)0.286500:0.000000,(Lama_glama:0.000000,PA12:0.000000,PA11:0.000000,(Bos_taurus:0.000000,PA15:0.000000,PA14:0.000000,PA13:0.000000)0.197333:0.000000)0.176000:0.000000)0.325600:0.000000)0.341250:0.000000)0.355059:0.000000)0.367556:0.000000,(Thomashuxleya_externa_comb:0.000000,(Carodnia_vieirai:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.144000:0.000000)0.362000:0.000000)0.397333:0.000000)0.324000:0.000000)0.301739:0.000000)0.290167:0.000000)0.280160:0.000000)0.271538:0.000000,(PA43:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000)0.044000:0.000000)0.022000:0.000000)0.235333:0.000000)0.220625:0.000000,(Orycteropus_afer:0.000000,((Dasypus_novemcinctus:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.792000:0.000000)0.584000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.508000:0.000000)0.401000:0.000000)0.352800:0.000000)0.195053:0.000000)0.154077:0.000000,(PA26:0.000000,(PA27:0.000000,(PA28:0.000000,((Rattus_norvegicus:0.000000,(Ictidomys_tridecemlineatus:0.000000,(PA30:0.000000,PA31:0.000000,PA29:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.616000:0.000000)0.402000:0.000000)0.360000:0.000000)0.341333:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.312000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.800000:0.000000)1.000000:0.000000)0.810667:0.000000)0.753000:0.000000)0.524000:0.000000)0.903385:0.000000)0.974857:0.000000)0.910667:0.000000,((PA33:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000,PA34:0.000000)1.000000:0.000000)0.698667:0.000000,((Notharctus_tenebrosus:0.000000,((Tarsius_syrichta:0.000000,PA36:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000,PA37:0.000000)0.996000:0.000000)0.755000:0.000000,(Lemur_catta:0.000000,PA35:0.000000)0.240000:0.000000)0.866667:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,(Tupaia_glis:0.000000,PA32:0.000000)0.032000:0.000000)1.000000:0.000000)0.718000:0.000000)0.649714:0.000000)0.482452:0.000000)0.588667:0.000000)0.623059:0.000000)0.746286:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.936000:0.000000)0.994667:0.000000)0.765600:0.000000,(Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.796000:0.000000)0.594000:0.000000)0.890286:0.000000)0.978667:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.828000:0.000000)0.972000:0.000000)1.000000:0.000000)1.000000:0.000000));

[“dental only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: 2 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(((((Leptictis_dakotensis,((Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))),(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(Tribosphenomys_minutus,(((Rattus_norvegicus,(Ictidomys_tridecemlineatus,(PA28,(Castor_canadensis,Cavia_porcellus)))),(Cocomys_lingchaensis,Paramys_delicatus)),(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))),(((PA40,((PA41,(PA42,(Procavia_capensis,((Mesohippus_bairdi,(Equus_caballus,(PA10,((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama))))),(Thomashuxleya_externa_comb,(Carodnia_vieirai,(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus)))))))),(PA43,(Rhynchocyon_cirnei,PA44)))),(PA39,PA38)),(Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))),((PA6,(((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))),(PA27,PA26))),(PA7,(PA8,(PA21,(PA9,(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))))),(PA5,(PA3,PA1)))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(((((Leptictis_dakotensis,((Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))),(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(Tribosphenomys_minutus,(((Rattus_norvegicus,(Ictidomys_tridecemlineatus,(PA28,(Castor_canadensis,Cavia_porcellus)))),(Cocomys_lingchaensis,Paramys_delicatus)),(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))),(((PA43,((PA41,(PA42,(Procavia_capensis,((Mesohippus_bairdi,(Equus_caballus,(PA10,((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama))))),(Thomashuxleya_externa_comb,(Carodnia_vieirai,(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus)))))))),(PA40,(PA39,PA38)))),(Rhynchocyon_cirnei,PA44)),(Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))),((PA6,(PA27,(PA26,((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))),(PA7,(PA8,(PA21,(PA9,(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))))),(PA5,(PA3,PA1)))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));

[“dental only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: strict consensus of 2 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.988000:0.000000,((Eomaia_scansoria:0.000000,((((Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.444000:0.000000)0.808000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.796000:0.000000)0.168000:0.000000)0.436000:0.000000,(((((Leptictis_dakotensis:0.000000,((Protungulatum_donnae:0.000000,(Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.304000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.348000:0.000000)0.104000:0.000000)0.216000:0.000000)0.100000:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)0.820000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.636000:0.000000)0.820000:0.000000)0.040000:0.000000)0.008000:0.000000,(Tribosphenomys_minutus:0.000000,(((Rattus_norvegicus:0.000000,(Ictidomys_tridecemlineatus:0.000000,(PA28:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.872000:0.000000)0.124000:0.000000)0.072000:0.000000)0.052000:0.000000,(Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.228000:0.000000)0.032000:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.788000:0.000000)0.988000:0.000000)0.428000:0.000000)0.004000:0.000000)0.844000:0.000000)0.000000:0.000000,((PA43:0.000000,PA40:0.000000,PA39:0.000000,PA38:0.000000,(PA41:0.000000,(PA42:0.000000,(Procavia_capensis:0.000000,((Mesohippus_bairdi:0.000000,(Equus_caballus:0.000000,(PA10:0.000000,((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.948000:0.000000)0.112000:0.000000)0.568000:0.000000)0.064000:0.000000)0.004000:0.000000)0.012000:0.000000)0.024000:0.000000)0.008000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.928000:0.000000)0.004000:0.000000)0.000000:0.000000)0.004000:0.000000)0.200000:0.000000,(Thomashuxleya_externa_comb:0.000000,(Carodnia_vieirai:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.152000:0.000000)0.260000:0.000000)0.308000:0.000000)0.056000:0.000000)0.024000:0.000000)0.012000:0.000000)0.000000:0.000000)0.012000:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000)0.336000:0.000000)0.000000:0.000000,(Orycteropus_afer:0.000000,((Dasypus_novemcinctus:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.768000:0.000000)0.380000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.488000:0.000000)0.128000:0.000000)0.120000:0.000000)0.000000:0.000000)0.000000:0.000000,((PA6:0.000000,(PA27:0.000000,PA26:0.000000,((PA33:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000)0.168000:0.000000,((Notharctus_tenebrosus:0.000000,(Lemur_catta:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.976000:0.000000)0.376000:0.000000)0.156000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.108000:0.000000)0.076000:0.000000)0.260000:0.000000)0.000000:0.000000,(PA7:0.000000,(PA21:0.000000,PA8:0.000000,(PA9:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.704000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.740000:0.000000)0.980000:0.000000)1.000000:0.000000)1.000000:0.000000)1.000000:0.000000)0.168000:0.000000)0.092000:0.000000)0.024000:0.000000)0.000000:0.000000)0.000000:0.000000,(PA5:0.000000,(PA3:0.000000,PA1:0.000000)0.128000:0.000000)0.000000:0.000000)0.000000:0.000000)0.508000:0.000000,(Dromiciops_gliroides:0.000000,(PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.952000:0.000000)0.028000:0.000000)0.996000:0.000000)0.372000:0.000000,(Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.824000:0.000000)0.304000:0.000000)0.344000:0.000000)0.988000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.828000:0.000000)0.952000:0.000000)1.000000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“dental only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: strict consensus of 2 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.988000:0.000000,((Eomaia_scansoria:0.000000,((((Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.444000:0.000000)0.880000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.796000:0.000000)0.643000:0.000000)0.684800:0.000000,(((((Leptictis_dakotensis:0.000000,((Protungulatum_donnae:0.000000,(Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.304000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.348000:0.000000)0.353333:0.000000)0.410000:0.000000)0.364000:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)0.820000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.818667:0.000000)0.866000:0.000000)0.401200:0.000000)0.366182:0.000000,(Tribosphenomys_minutus:0.000000,(((Rattus_norvegicus:0.000000,(Ictidomys_tridecemlineatus:0.000000,(PA28:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.872000:0.000000)0.498000:0.000000)0.382667:0.000000)0.306000:0.000000,(Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.228000:0.000000)0.278000:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.788000:0.000000)0.988000:0.000000)0.801333:0.000000)0.816400:0.000000)0.906909:0.000000)0.440000:0.000000,((PA43:0.000000,PA40:0.000000,PA39:0.000000,PA38:0.000000,(PA41:0.000000,(PA42:0.000000,(Procavia_capensis:0.000000,((Mesohippus_bairdi:0.000000,(Equus_caballus:0.000000,(PA10:0.000000,((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.948000:0.000000)0.532000:0.000000)0.717333:0.000000)0.555000:0.000000)0.447200:0.000000)0.381333:0.000000)0.338286:0.000000)0.303000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.928000:0.000000)0.324400:0.000000)0.344727:0.000000)0.369667:0.000000)0.395692:0.000000,(Thomashuxleya_externa_comb:0.000000,(Carodnia_vieirai:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.152000:0.000000)0.384000:0.000000)0.418667:0.000000)0.341000:0.000000)0.313333:0.000000)0.298947:0.000000)0.285800:0.000000)0.274857:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000)0.336000:0.000000)0.213778:0.000000,(Orycteropus_afer:0.000000,((Dasypus_novemcinctus:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.768000:0.000000)0.578000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.488000:0.000000)0.401000:0.000000)0.360000:0.000000)0.183394:0.000000)0.240667:0.000000,((PA6:0.000000,(PA27:0.000000,PA26:0.000000,((PA33:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000)0.584000:0.000000,((Notharctus_tenebrosus:0.000000,(Lemur_catta:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.976000:0.000000)0.676000:0.000000)0.718667:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.702667:0.000000)0.588444:0.000000)0.576727:0.000000)0.528667:0.000000,(PA7:0.000000,(PA21:0.000000,PA8:0.000000,(PA9:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.704000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.740000:0.000000)0.984000:0.000000)1.000000:0.000000)1.000000:0.000000)1.000000:0.000000)0.896000:0.000000)0.735600:0.000000)0.670909:0.000000)0.349333:0.000000)0.449565:0.000000,(PA5:0.000000,(PA3:0.000000,PA1:0.000000)0.128000:0.000000)0.064000:0.000000)0.515600:0.000000)0.734286:0.000000,(Dromiciops_gliroides:0.000000,(PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.952000:0.000000)0.514000:0.000000)0.997333:0.000000)0.764800:0.000000,(Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.824000:0.000000)0.590000:0.000000)0.899429:0.000000)0.992000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.828000:0.000000)0.976000:0.000000)1.000000:0.000000)1.000000:0.000000)-1.#IND00:0.000000);

[“dental only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: 12 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(PA7,(((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(((Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))),(PA38,(PA40,PA39))),(Leptictis_dakotensis,Rhynchocyon_cirnei))),((PA6,(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))),(PA5,(PA3,PA1)))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((PA5,(PA6,((PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus)))),(((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(((PA38,(Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))),(PA40,PA39)),(Leptictis_dakotensis,Rhynchocyon_cirnei))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((PA5,(PA6,((PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus)))),((((Leptictis_dakotensis,((Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))),(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(Tribosphenomys_minutus,((Paramys_delicatus,(Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus)))),(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))),((Rhynchocyon_cirnei,(Procavia_capensis,((((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)),(Thomashuxleya_externa_comb,(Carodnia_vieirai,(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))))))),((Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))),(PA39,(PA40,PA38))))),((PA26,((Cynocephalus_volans,Galeopterus_variegatus),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((PA5,(PA6,((PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus)))),(((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(((((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)),(Orycteropus_afer,PA39)),(PA40,PA38)),(Leptictis_dakotensis,Rhynchocyon_cirnei))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))))),(PA3,PA1)),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((PA3,(PA1,(PA5,(PA6,((PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus)))),(((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(((Orycteropus_afer,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))),(PA40,PA38)),(Leptictis_dakotensis,Rhynchocyon_cirnei))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((PA5,(PA6,((PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus)))),(((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(((PA39,(Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))),(PA40,PA38)),(Leptictis_dakotensis,Rhynchocyon_cirnei))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((PA1,(PA3,(PA5,((PA6,(PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))))),(((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),((PA40,(PA38,(Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,(Tamandua_tetradactyla,PA39))),(Manis_pentadactyla,Metacheiromys_marshi))))),(Leptictis_dakotensis,Rhynchocyon_cirnei))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((PA3,(PA1,(PA5,((PA6,(PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))))),(((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(((Orycteropus_afer,((PA39,(Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla))),(Manis_pentadactyla,Metacheiromys_marshi))),(PA40,PA38)),(Leptictis_dakotensis,Rhynchocyon_cirnei))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((PA3,(PA1,(PA5,(PA6,((PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus)))),(((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),((PA40,(PA38,(Orycteropus_afer,((Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla))),(Manis_pentadactyla,Metacheiromys_marshi))))),(Leptictis_dakotensis,Rhynchocyon_cirnei))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((PA5,((PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus)))),(PA6,(((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),((PA38,(PA40,(Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,(Metacheiromys_marshi,PA39)))))),(Leptictis_dakotensis,Rhynchocyon_cirnei))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))))),(PA3,PA1)),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((PA1,(PA3,(PA5,(PA6,((PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus)))),(((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),((PA38,(PA40,(Orycteropus_afer,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(PA39,(Manis_pentadactyla,Metacheiromys_marshi)))))),(Leptictis_dakotensis,Rhynchocyon_cirnei))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((PA5,(PA6,((PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus)))),(((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(((Orycteropus_afer,(PA39,((Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))),(PA40,PA38)),(Leptictis_dakotensis,Rhynchocyon_cirnei))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));

[“dental only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: strict consensus of 12 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.996000:0.000000,((Eomaia_scansoria:0.000000,(((PA7:0.000000,PA6:0.000000,PA5:0.000000,PA3:0.000000,PA1:0.000000,(Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.456000:0.000000)0.824000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.868000:0.000000)0.208000:0.000000)0.448000:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000,(Protungulatum_donnae:0.000000,(Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.244000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.352000:0.000000)0.080000:0.000000)0.272000:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)0.996000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.736000:0.000000)1.000000:0.000000)0.012000:0.000000,(Procavia_capensis:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.976000:0.000000)0.096000:0.000000)0.556000:0.000000)0.052000:0.000000)0.016000:0.000000)0.032000:0.000000)0.036000:0.000000)0.012000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.972000:0.000000)0.016000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.552000:0.000000)0.216000:0.000000,(Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000,(Loxodonta_africana:0.000000,Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.340000:0.000000)0.032000:0.000000)0.140000:0.000000,(Paramys_delicatus:0.000000,Tribosphenomys_minutus:0.000000,(Rattus_norvegicus:0.000000,((Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.848000:0.000000,(Cocomys_lingchaensis:0.000000,Ictidomys_tridecemlineatus:0.000000)0.360000:0.000000)0.216000:0.000000)0.108000:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.860000:0.000000)1.000000:0.000000)0.440000:0.000000)0.992000:0.000000,(Dasypus_novemcinctus:0.000000,Hapalops_elongatus:0.000000,Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000,Orycteropus_afer:0.000000,Tamandua_tetradactyla:0.000000,PA40:0.000000,PA39:0.000000,PA38:0.000000)0.012000:0.000000)0.000000:0.000000,(PA26:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,((Notharctus_tenebrosus:0.000000,(Lemur_catta:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.988000:0.000000)0.424000:0.000000)0.204000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.124000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)0.996000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.692000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.756000:0.000000)0.964000:0.000000)1.000000:0.000000)0.996000:0.000000)1.000000:0.000000)0.060000:0.000000)0.476000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.952000:0.000000)0.996000:0.000000)0.404000:0.000000,(Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.816000:0.000000)0.320000:0.000000)0.300000:0.000000)0.996000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.828000:0.000000)0.940000:0.000000)1.000000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“dental only” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: strict consensus of 12 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.996000:0.000000,((Eomaia_scansoria:0.000000,(((PA7:0.000000,PA6:0.000000,PA5:0.000000,PA3:0.000000,PA1:0.000000,(Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.456000:0.000000)0.876000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.868000:0.000000)0.644000:0.000000)0.674400:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000,(Protungulatum_donnae:0.000000,(Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.244000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.352000:0.000000)0.341333:0.000000)0.410000:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)0.996000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.912000:0.000000)1.000000:0.000000)0.414800:0.000000,(Procavia_capensis:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.976000:0.000000)0.538000:0.000000)0.720000:0.000000)0.553000:0.000000)0.452000:0.000000)0.392000:0.000000)0.348000:0.000000)0.330500:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.972000:0.000000)0.356400:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.552000:0.000000)0.418000:0.000000,(Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000,(Loxodonta_africana:0.000000,Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.522000:0.000000)0.480000:0.000000)0.383333:0.000000,(Paramys_delicatus:0.000000,Tribosphenomys_minutus:0.000000,(Rattus_norvegicus:0.000000,((Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.848000:0.000000,(Cocomys_lingchaensis:0.000000,Ictidomys_tridecemlineatus:0.000000)0.360000:0.000000)0.549333:0.000000)0.524000:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.860000:0.000000)1.000000:0.000000)0.813333:0.000000)0.994400:0.000000,(Dasypus_novemcinctus:0.000000,Hapalops_elongatus:0.000000,Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000,Orycteropus_afer:0.000000,Tamandua_tetradactyla:0.000000,PA40:0.000000,PA39:0.000000,PA38:0.000000)0.184000:0.000000)0.314419:0.000000,(PA26:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,((Notharctus_tenebrosus:0.000000,(Lemur_catta:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.988000:0.000000)0.706000:0.000000)0.734667:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.708000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)0.996000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.692000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.756000:0.000000)0.974000:0.000000)1.000000:0.000000)0.999333:0.000000)1.000000:0.000000)0.500706:0.000000)0.730857:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.952000:0.000000)0.997333:0.000000)0.777200:0.000000,(Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.816000:0.000000)0.606000:0.000000)0.897714:0.000000)0.997333:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.828000:0.000000)0.970000:0.000000)1.000000:0.000000)1.000000:0.000000)-1.#IND00:0.000000);

[“max preservation" PAs corrected for the O'Leary et al. character ontology: 24 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA2,(Tachyglossus_aculeatus,PA1))),(((PA3,((((PA43,(Orycteropus_afer,((PA44,(Echinops_telfairi,(Amblysomus_hottentotus,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)))),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA5,(((PA8,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12)),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA7,(((Erinaceus_europaeus,PA24),(Talpa_europaea,(Sorex_araneus,PA25))),(Solenodon_paradoxus,PA23)))),(PA6,((PA28,((PA29,((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(((PA33,(PA34,(Cynocephalus_volans,Galeopterus_variegatus))),((PA35,(Tarsius_syrichta,(PA36,(PA37,(Homo_sapiens,Saimiri_sciureus))))),(Lemur_catta,Notharctus_tenebrosus))),(PA26,(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,(PA5,((((PA40,(Orycteropus_afer,(PA43,((PA44,(Echinops_telfairi,(Amblysomus_hottentotus,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),((PA6,(((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA7,(Solenodon_paradoxus,(PA23,((Erinaceus_europaeus,PA24),(Sorex_araneus,(Talpa_europaea,PA25)))))))),((PA28,((PA29,((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(((PA33,(Cynocephalus_volans,(Galeopterus_variegatus,PA34))),((PA35,((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36))),(Lemur_catta,Notharctus_tenebrosus))),(PA26,(PA27,(PA32,(Ptilocercus_lowii,Tupaia_glis))))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,(PA5,(((PA40,((((PA44,(Echinops_telfairi,(Amblysomus_hottentotus,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(Orycteropus_afer,PA43)),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),((PA6,((PA7,(PA8,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(PA11,((Sus_scrofa,(PA12,(PA13,(Hippopotamus_amphibius,(PA14,((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15)))))))),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),((Erinaceus_europaeus,(PA24,(Sorex_araneus,(Talpa_europaea,PA25)))),(Solenodon_paradoxus,PA23)))),((PA28,(((PA29,(PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA26,(((PA33,(Cynocephalus_volans,(Galeopterus_variegatus,PA34))),((PA35,((PA37,(Homo_sapiens,Saimiri_sciureus)),(Tarsius_syrichta,PA36))),(Lemur_catta,Notharctus_tenebrosus))),(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,(((PA38,(PA40,((((PA44,(Amblysomus_hottentotus,(Echinops_telfairi,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(Orycteropus_afer,PA43)),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),(PA39,(Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA5,((PA6,((PA7,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((Sus_scrofa,(PA12,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus))),(Hippopotamus_amphibius,PA14))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA23,(Solenodon_paradoxus,(Erinaceus_europaeus,(PA24,(Talpa_europaea,(Sorex_araneus,PA25)))))))),((PA28,((PA29,((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,((PA33,(Galeopterus_variegatus,(Cynocephalus_volans,PA34))),(((Homo_sapiens,(Saimiri_sciureus,PA37)),(Tarsius_syrichta,PA36)),(PA35,(Lemur_catta,Notharctus_tenebrosus))))),(PA27,(Tupaia_glis,(Ptilocercus_lowii,PA32)))))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,(PA2,PA1))),(((PA3,((((PA40,(((PA44,(Amblysomus_hottentotus,(Echinops_telfairi,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(Orycteropus_afer,PA43))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA5,(((PA7,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA12,(Sus_scrofa,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),((Erinaceus_europaeus,(PA24,(Sorex_araneus,(Talpa_europaea,PA25)))),(Solenodon_paradoxus,PA23))),(PA6,((PA28,((PA29,((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,(PA33,((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),((PA35,((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36))),(Lemur_catta,Notharctus_tenebrosus))))),(PA27,(Tupaia_glis,(Ptilocercus_lowii,PA32)))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,(((PA38,(PA40,((((PA44,(Echinops_telfairi,(Amblysomus_hottentotus,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(Orycteropus_afer,PA43)),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))),(PA5,((PA6,((PA7,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA12,(Sus_scrofa,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA23,(Solenodon_paradoxus,(Erinaceus_europaeus,(PA24,(PA25,(Sorex_araneus,Talpa_europaea)))))))),((PA28,(((PA29,((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,((PA33,(Galeopterus_variegatus,(Cynocephalus_volans,PA34))),(PA35,(((Saimiri_sciureus,(Homo_sapiens,PA37)),(Tarsius_syrichta,PA36)),(Lemur_catta,Notharctus_tenebrosus))))),(PA27,(PA32,(Ptilocercus_lowii,Tupaia_glis)))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,(PA2,PA1))),(((PA3,(((PA40,((((PA44,(Amblysomus_hottentotus,(Echinops_telfairi,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(Orycteropus_afer,PA43)),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA5,(((PA7,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA12,(Sus_scrofa,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus))),(Hippopotamus_amphibius,PA14))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA23,(Solenodon_paradoxus,(Erinaceus_europaeus,(PA24,(Sorex_araneus,(Talpa_europaea,PA25))))))),(PA6,((PA28,((PA29,((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA26,((PA33,((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),((PA35,((Homo_sapiens,(Saimiri_sciureus,PA37)),(Tarsius_syrichta,PA36))),(Lemur_catta,Notharctus_tenebrosus)))),(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,((PA5,((PA38,((Orycteropus_afer,(PA43,((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA6,((PA7,(((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA12,(Sus_scrofa,(PA13,(Hippopotamus_amphibius,(PA14,((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus)))))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),(Solenodon_paradoxus,(PA23,((Erinaceus_europaeus,PA24),(Talpa_europaea,(Sorex_araneus,PA25))))))),((PA28,(((PA29,((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(((PA33,(Cynocephalus_volans,(Galeopterus_variegatus,PA34))),((Tarsius_syrichta,(PA36,(PA37,(Homo_sapiens,Saimiri_sciureus)))),(PA35,(Lemur_catta,Notharctus_tenebrosus)))),(PA26,(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,(PA5,((PA38,((((PA44,((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(Leptictis_dakotensis,Rhynchocyon_cirnei))),(Orycteropus_afer,PA43)),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA6,((PA7,((PA8,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA12,(Sus_scrofa,(PA13,(Hippopotamus_amphibius,(PA14,((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15)))))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(((Erinaceus_europaeus,PA24),(Sorex_araneus,(Talpa_europaea,PA25))),(Solenodon_paradoxus,PA23)))),((PA28,((PA29,(((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(PA33,((Tarsius_syrichta,(PA36,(PA37,(Homo_sapiens,Saimiri_sciureus)))),(PA35,(Lemur_catta,Notharctus_tenebrosus))))),(PA26,(PA27,(PA32,(Ptilocercus_lowii,Tupaia_glis))))))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,(((PA40,((((PA44,(Echinops_telfairi,(Amblysomus_hottentotus,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(Orycteropus_afer,PA43)),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA5,((PA6,(PA7,((PA8,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(((PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus)))))),(Sus_scrofa,PA12)),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),((Erinaceus_europaeus,(PA24,(PA25,(Sorex_araneus,Talpa_europaea)))),(Solenodon_paradoxus,PA23))))),((PA28,(((PA29,(Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus)))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA26,((PA33,((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(((Homo_sapiens,(Saimiri_sciureus,PA37)),(Tarsius_syrichta,PA36)),(PA35,(Lemur_catta,Notharctus_tenebrosus))))),(PA27,(PA32,(Ptilocercus_lowii,Tupaia_glis)))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),((((((PA38,(PA40,((PA43,(Orycteropus_afer,((PA44,(PA45,(Amblysomus_hottentotus,Echinops_telfairi))),(Leptictis_dakotensis,Rhynchocyon_cirnei)))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))),(PA5,(PA6,((((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(PA11,(((PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15)))))),(Sus_scrofa,PA12)),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),(PA7,((Erinaceus_europaeus,(PA24,(Sorex_araneus,(Talpa_europaea,PA25)))),(Solenodon_paradoxus,PA23)))),((PA28,(((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA26,(((PA33,(Galeopterus_variegatus,(Cynocephalus_volans,PA34))),((Tarsius_syrichta,(PA36,(Homo_sapiens,(Saimiri_sciureus,PA37)))),(PA35,(Lemur_catta,Notharctus_tenebrosus)))),(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))),(PA3,PA1)),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,(PA5,((((((PA44,(Echinops_telfairi,(Amblysomus_hottentotus,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(Orycteropus_afer,PA43)),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA6,(((PA7,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,(PA12,(Sus_scrofa,(PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus))))))))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),((Erinaceus_europaeus,(PA24,(PA25,(Sorex_araneus,Talpa_europaea)))),(Solenodon_paradoxus,PA23))),((PA28,((PA29,(((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA27,((PA26,((PA33,(Cynocephalus_volans,(Galeopterus_variegatus,PA34))),(PA35,((PA36,(Tarsius_syrichta,(PA37,(Homo_sapiens,Saimiri_sciureus)))),(Lemur_catta,Notharctus_tenebrosus))))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,(PA5,((((PA43,(Orycteropus_afer,((PA44,(Amblysomus_hottentotus,(Echinops_telfairi,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)))),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA6,(((PA7,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA11,(PA12,(Sus_scrofa,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14)))))),(Bos_taurus,Lama_glama))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),((Erinaceus_europaeus,(PA24,(PA25,(Sorex_araneus,Talpa_europaea)))),(Solenodon_paradoxus,PA23))),((PA28,((PA29,(((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA27,((PA26,((PA33,(Galeopterus_variegatus,(Cynocephalus_volans,PA34))),(PA35,((Tarsius_syrichta,(PA36,(PA37,(Homo_sapiens,Saimiri_sciureus)))),(Lemur_catta,Notharctus_tenebrosus))))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,((((((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))),(Orycteropus_afer,PA43)),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA5,((PA6,(((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(PA11,((PA12,(Sus_scrofa,(PA13,(Hippopotamus_amphibius,(PA14,((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA7,(PA23,(Solenodon_paradoxus,(Erinaceus_europaeus,(PA24,(Sorex_araneus,(Talpa_europaea,PA25))))))))),((PA28,(((PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus))),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA27,((PA26,((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(PA33,(((Homo_sapiens,(Saimiri_sciureus,PA37)),(Tarsius_syrichta,PA36)),(PA35,(Lemur_catta,Notharctus_tenebrosus)))))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,((((Orycteropus_afer,(PA43,((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA5,((PA6,(((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(PA11,((PA12,(Sus_scrofa,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))))),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA7,(PA23,(Solenodon_paradoxus,(Erinaceus_europaeus,(PA24,(Talpa_europaea,(Sorex_araneus,PA25))))))))),((PA28,((PA29,((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA27,((PA26,((PA33,(PA34,(Cynocephalus_volans,Galeopterus_variegatus))),(((Homo_sapiens,(Saimiri_sciureus,PA37)),(Tarsius_syrichta,PA36)),(PA35,(Lemur_catta,Notharctus_tenebrosus))))),(Tupaia_glis,(Ptilocercus_lowii,PA32)))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,((PA5,((PA38,((((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))),(Orycteropus_afer,PA43)),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),((PA6,(((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12)),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),(PA7,(((Erinaceus_europaeus,PA24),(Talpa_europaea,(Sorex_araneus,PA25))),(Solenodon_paradoxus,PA23))))),((PA28,(((PA29,((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA27,(PA26,(PA33,((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),(PA35,((PA36,(Tarsius_syrichta,(Homo_sapiens,(Saimiri_sciureus,PA37)))),(Lemur_catta,Notharctus_tenebrosus))))))),(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),((((PA5,((((PA40,(PA43,(Orycteropus_afer,((PA44,(Echinops_telfairi,(Amblysomus_hottentotus,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(PA38,(PA39,(Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA6,(((PA7,(PA8,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12)),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(Solenodon_paradoxus,(PA23,((Erinaceus_europaeus,PA24),(Sorex_araneus,(Talpa_europaea,PA25)))))),((PA28,(((PA29,(PA31,(Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(PA33,(PA35,((Tarsius_syrichta,(PA36,(Saimiri_sciureus,(Homo_sapiens,PA37)))),(Lemur_catta,Notharctus_tenebrosus)))))),(PA27,(Tupaia_glis,(Ptilocercus_lowii,PA32))))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA2,(Tachyglossus_aculeatus,PA1))),(((PA3,(PA5,((((PA43,(Orycteropus_afer,((PA44,(PA45,(Amblysomus_hottentotus,Echinops_telfairi))),(Leptictis_dakotensis,Rhynchocyon_cirnei)))),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA6,((((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(PA11,(((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12)),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA7,(Solenodon_paradoxus,(PA23,((Erinaceus_europaeus,PA24),(Sorex_araneus,(Talpa_europaea,PA25))))))),((PA28,((PA29,(((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,(PA33,((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(((PA37,(Homo_sapiens,Saimiri_sciureus)),(Tarsius_syrichta,PA36)),(PA35,(Lemur_catta,Notharctus_tenebrosus)))))),(PA27,(PA32,(Ptilocercus_lowii,Tupaia_glis)))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA2,(Tachyglossus_aculeatus,PA1))),(((PA3,((((PA40,(((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei))),(Orycteropus_afer,PA43))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(PA38,((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA5,(PA6,(((PA7,(PA8,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(PA11,((Sus_scrofa,(PA12,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15))),(Hippopotamus_amphibius,PA14))))),(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA23,(Solenodon_paradoxus,((Erinaceus_europaeus,PA24),(PA25,(Sorex_araneus,Talpa_europaea)))))),((PA28,(((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(PA33,(((Homo_sapiens,(Saimiri_sciureus,PA37)),(Tarsius_syrichta,PA36)),(PA35,(Lemur_catta,Notharctus_tenebrosus)))))),(PA27,(Ptilocercus_lowii,(Tupaia_glis,PA32)))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),(((PA1,(PA3,((PA38,(((PA40,(Orycteropus_afer,(PA43,(PA44,((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),(Leptictis_dakotensis,Rhynchocyon_cirnei)))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(PA39,(Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA5,((PA6,(((PA8,(((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((Sus_scrofa,(PA12,(PA13,(Hippopotamus_amphibius,(PA14,((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15)))))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20)))))),(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))),(PA7,(PA23,(Solenodon_paradoxus,((Erinaceus_europaeus,PA24),(PA25,(Sorex_araneus,Talpa_europaea)))))))),((PA28,((PA29,((Ictidomys_tridecemlineatus,(PA31,(Castor_canadensis,Cavia_porcellus))),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),(PA33,(PA35,((Tarsius_syrichta,(PA36,(Homo_sapiens,(Saimiri_sciureus,PA37)))),(Lemur_catta,Notharctus_tenebrosus)))))),(PA27,(PA32,(Ptilocercus_lowii,Tupaia_glis)))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,((PA5,((PA38,((Orycteropus_afer,(PA43,(PA44,((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(Leptictis_dakotensis,Rhynchocyon_cirnei))))),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),((PA7,(((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((Sus_scrofa,(PA12,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15))),(Hippopotamus_amphibius,PA14))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),((Erinaceus_europaeus,(PA24,(PA25,(Sorex_araneus,Talpa_europaea)))),(Solenodon_paradoxus,PA23)))),(PA6,((PA28,((PA29,(((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA26,(PA33,((PA34,(Cynocephalus_volans,Galeopterus_variegatus)),((Tarsius_syrichta,(PA36,(Homo_sapiens,(Saimiri_sciureus,PA37)))),(PA35,(Lemur_catta,Notharctus_tenebrosus)))))),(PA27,(PA32,(Ptilocercus_lowii,Tupaia_glis))))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(Tachyglossus_aculeatus,PA2)),((((PA5,(((PA38,((((PA44,(Echinops_telfairi,(Amblysomus_hottentotus,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(Orycteropus_afer,PA43)),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))),(PA6,(((PA7,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((PA12,(Sus_scrofa,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Tursiops_truncatus,(Caperea_marginata,PA15))),(Hippopotamus_amphibius,PA14))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA23,(Solenodon_paradoxus,((Erinaceus_europaeus,PA24),(Sorex_araneus,(Talpa_europaea,PA25)))))),((PA28,(((PA29,((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),(PA27,((PA26,((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(PA33,((PA35,((PA37,(Homo_sapiens,Saimiri_sciureus)),(Tarsius_syrichta,PA36))),(Lemur_catta,Notharctus_tenebrosus))))),(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))),(PA3,PA1)),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,(((PA38,((PA43,(Orycteropus_afer,((PA44,(Amblysomus_hottentotus,(Echinops_telfairi,PA45))),(Leptictis_dakotensis,Rhynchocyon_cirnei)))),(PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))))),(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA5,(PA6,((((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(((PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(Caperea_marginata,(Tursiops_truncatus,PA15))),(Hippopotamus_amphibius,PA14))),(Sus_scrofa,PA12)),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA8,(PA21,(((Felis_silvestris,(Canis_lupus,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA7,(((Erinaceus_europaeus,PA24),(Talpa_europaea,(Sorex_araneus,PA25))),(Solenodon_paradoxus,PA23)))),((PA28,(((PA29,((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA27,(PA26,((PA33,(PA34,(Cynocephalus_volans,Galeopterus_variegatus))),((PA35,(Tarsius_syrichta,(PA36,(Homo_sapiens,(Saimiri_sciureus,PA37))))),(Lemur_catta,Notharctus_tenebrosus))))),(Ptilocercus_lowii,(Tupaia_glis,PA32))))))))),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Ornithorhynchus_anatinus,(PA1,(Tachyglossus_aculeatus,PA2))),(((PA3,((((PA40,(Orycteropus_afer,(PA43,((Amblysomus_hottentotus,(Echinops_telfairi,PA45)),(PA44,(Leptictis_dakotensis,Rhynchocyon_cirnei)))))),(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(PA38,(PA39,(Dasypus_novemcinctus,(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA5,(PA6,(((PA8,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,(((PA13,(PA14,(Hippopotamus_amphibius,((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus)))))),(Sus_scrofa,PA12)),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,(((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi))))),(PA7,(((Erinaceus_europaeus,PA24),(Sorex_araneus,(Talpa_europaea,PA25))),(Solenodon_paradoxus,PA23)))),((PA28,(((PA29,((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31))),(Rattus_norvegicus,PA30)),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))),((PA27,(PA26,((PA33,(Galeopterus_variegatus,(Cynocephalus_volans,PA34))),((Tarsius_syrichta,(PA36,(Saimiri_sciureus,(Homo_sapiens,PA37)))),(PA35,(Lemur_catta,Notharctus_tenebrosus)))))),(PA32,(Ptilocercus_lowii,Tupaia_glis))))))))),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));

[“max preservation” PAs corrected for the O'Leary et al. character ontology: strict consensus of 24 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000,PA3:0.000000,PA2:0.000000,PA1:0.000000,(PA5:0.000000,(PA38:0.000000,(PA40:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(PA44:0.000000,(Amblysomus_hottentotus:0.000000,Echinops_telfairi:0.000000,PA45:0.000000)0.440000:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.020000:0.000000)0.012000:0.000000)0.008000:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.256000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.116000:0.000000)0.088000:0.000000)0.032000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.288000:0.000000)0.016000:0.000000)0.072000:0.000000)0.000000:0.000000,(Dasypus_novemcinctus:0.000000,PA39:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.988000:0.000000)0.604000:0.000000)0.000000:0.000000,(PA6:0.000000,(PA7:0.000000,(PA8:0.000000,(((((Apheliscus_insidiosus:0.000000,Protungulatum_donnae:0.000000)0.020000:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.324000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.272000:0.000000)0.040000:0.000000)0.164000:0.000000,(Rodhocetus_balochistanensis:0.000000,(Archaeotherium_mortoni:0.000000,Mesonyx_obtusidens:0.000000)0.040000:0.000000)0.056000:0.000000)0.000000:0.000000,((PA10:0.000000,(PA11:0.000000,(Sus_scrofa:0.000000,PA12:0.000000,(PA13:0.000000,(Hippopotamus_amphibius:0.000000,PA14:0.000000,((Artiocetus_clavis:0.000000,Basilosaurus_cetoides:0.000000)0.236000:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000,PA15:0.000000)0.992000:0.000000)0.156000:0.000000)0.148000:0.000000)0.004000:0.000000)0.004000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.024000:0.000000)0.024000:0.000000)0.004000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.348000:0.000000)0.016000:0.000000)0.004000:0.000000,(PA9:0.000000,(((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,(((Myotis_lucifugus:0.000000,PA17:0.000000)0.228000:0.000000,(Pteronotus_parnellii:0.000000,PA18:0.000000)0.096000:0.000000)0.400000:0.000000,((Nycteris_thebaica:0.000000,PA19:0.000000)0.064000:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.444000:0.000000)0.748000:0.000000)0.596000:0.000000)0.452000:0.000000,(PA16:0.000000,(Pteropus_giganteus:0.000000,PA20:0.000000)0.468000:0.000000)0.108000:0.000000)1.000000:0.000000)0.100000:0.000000)0.000000:0.000000,(PA21:0.000000,(((Canis_lupus:0.000000,Felis_silvestris:0.000000,PA22:0.000000)0.780000:0.000000,(Sinopa_rapax:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.928000:0.000000)0.956000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.964000:0.000000)0.068000:0.000000)0.184000:0.000000)0.000000:0.000000,(Solenodon_paradoxus:0.000000,PA23:0.000000,(Erinaceus_europaeus:0.000000,PA24:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000,PA25:0.000000)0.812000:0.000000)0.148000:0.000000)0.168000:0.000000)0.000000:0.000000,((PA28:0.000000,((PA29:0.000000,(Ictidomys_tridecemlineatus:0.000000,PA31:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.180000:0.000000)0.132000:0.000000,(Rattus_norvegicus:0.000000,PA30:0.000000)0.176000:0.000000)0.236000:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.600000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.736000:0.000000)0.984000:0.000000)0.404000:0.000000)0.356000:0.000000)0.344000:0.000000)0.552000:0.000000)0.996000:0.000000,(PA27:0.000000,PA26:0.000000,(PA33:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000,PA34:0.000000)1.000000:0.000000,(PA35:0.000000,(Tarsius_syrichta:0.000000,PA36:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000,PA37:0.000000)1.000000:0.000000)0.512000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000)0.368000:0.000000)1.000000:0.000000)0.796000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000,PA32:0.000000)1.000000:0.000000)0.508000:0.000000)0.736000:0.000000)0.000000:0.000000)0.128000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.972000:0.000000)0.968000:0.000000,((Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000)0.456000:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.344000:0.000000)0.584000:0.000000)0.584000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.980000:0.000000)1.000000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“max preservation” PAs corrected for the O'Leary et al. character ontology: strict consensus of 24 MPTs - nodal values = "transfer bootstrap expectation" values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000,PA3:0.000000,PA2:0.000000,PA1:0.000000,(PA5:0.000000,(PA38:0.000000,(PA40:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(PA44:0.000000,(Amblysomus_hottentotus:0.000000,Echinops_telfairi:0.000000,PA45:0.000000)0.468000:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.020000:0.000000)0.234400:0.000000)0.178286:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.256000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.116000:0.000000)0.200000:0.000000)0.236000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.288000:0.000000)0.269333:0.000000)0.292571:0.000000)0.168750:0.000000,(Dasypus_novemcinctus:0.000000,PA39:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.988000:0.000000)0.734667:0.000000)0.164000:0.000000,(PA6:0.000000,(PA7:0.000000,(PA8:0.000000,(((((Apheliscus_insidiosus:0.000000,Protungulatum_donnae:0.000000)0.020000:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.324000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.272000:0.000000)0.250667:0.000000)0.320800:0.000000,(Rodhocetus_balochistanensis:0.000000,(Archaeotherium_mortoni:0.000000,Mesonyx_obtusidens:0.000000)0.040000:0.000000)0.108000:0.000000)0.213500:0.000000,((PA10:0.000000,(PA11:0.000000,(Sus_scrofa:0.000000,PA12:0.000000,(PA13:0.000000,(Hippopotamus_amphibius:0.000000,PA14:0.000000,((Artiocetus_clavis:0.000000,Basilosaurus_cetoides:0.000000)0.236000:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000,PA15:0.000000)0.996000:0.000000)0.610000:0.000000)0.484000:0.000000)0.416000:0.000000)0.330667:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.024000:0.000000)0.359000:0.000000)0.371692:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.348000:0.000000)0.406133:0.000000)0.354833:0.000000,(PA9:0.000000,(((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,(((Myotis_lucifugus:0.000000,PA17:0.000000)0.228000:0.000000,(Pteronotus_parnellii:0.000000,PA18:0.000000)0.096000:0.000000)0.406667:0.000000,((Nycteris_thebaica:0.000000,PA19:0.000000)0.064000:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.444000:0.000000)0.796000:0.000000)0.768000:0.000000)0.841333:0.000000,(PA16:0.000000,(Pteropus_giganteus:0.000000,PA20:0.000000)0.468000:0.000000)0.288000:0.000000)1.000000:0.000000)0.930769:0.000000)0.380632:0.000000,(PA21:0.000000,(((Canis_lupus:0.000000,Felis_silvestris:0.000000,PA22:0.000000)0.780000:0.000000,(Sinopa_rapax:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.964000:0.000000)0.972800:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.964000:0.000000)0.730857:0.000000)0.699500:0.000000)0.302667:0.000000,(Solenodon_paradoxus:0.000000,PA23:0.000000,(Erinaceus_europaeus:0.000000,PA24:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000,PA25:0.000000)0.818000:0.000000)0.523000:0.000000)0.441333:0.000000)0.276571:0.000000,((PA28:0.000000,((PA29:0.000000,(Ictidomys_tridecemlineatus:0.000000,PA31:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.180000:0.000000)0.166667:0.000000,(Rattus_norvegicus:0.000000,PA30:0.000000)0.176000:0.000000)0.308000:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.600000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.736000:0.000000)0.986000:0.000000)0.797333:0.000000)0.748000:0.000000)0.661333:0.000000)0.962154:0.000000)0.996286:0.000000,(PA27:0.000000,PA26:0.000000,(PA33:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000,PA34:0.000000)1.000000:0.000000,(PA35:0.000000,(Tarsius_syrichta:0.000000,PA36:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000,PA37:0.000000)1.000000:0.000000)0.819000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000)0.368000:0.000000)1.000000:0.000000)0.926182:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000,PA32:0.000000)1.000000:0.000000)0.797000:0.000000)0.864516:0.000000)0.181400:0.000000)0.401176:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.972000:0.000000)0.969333:0.000000,((Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000)0.456000:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.344000:0.000000)0.673333:0.000000)0.793000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.980000:0.000000)1.000000:0.000000)1.000000:0.000000));

[“max preservation” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: 8 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((((Solenodon_paradoxus,(Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea))),(Echinops_telfairi,Erinaceus_europaeus)),(Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))))),(((PA7,(PA8,(((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),(PA21,(((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA6,(PA26,((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA33,(((Cynocephalus_volans,Galeopterus_variegatus),(Ptilocercus_lowii,Tupaia_glis)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus)))))))),(PA5,(PA38,(((PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(Orycteropus_afer,(PA43,(Rhynchocyon_cirnei,PA44)))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla))))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(PA7,(PA8,((PA21,(((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))),(Manis_pentadactyla,Metacheiromys_marshi))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))))))),(PA6,(PA26,((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))))),(PA5,(PA38,(((PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(PA43,(Orycteropus_afer,(Rhynchocyon_cirnei,PA44)))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((((Solenodon_paradoxus,(Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea))),(Echinops_telfairi,Erinaceus_europaeus)),(Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))))),(((PA7,(PA8,(((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),(PA21,(((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA6,(PA26,((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA33,(((Cynocephalus_volans,Galeopterus_variegatus),(Ptilocercus_lowii,Tupaia_glis)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus)))))))),(PA5,((PA38,(PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA43,(Orycteropus_afer,(Rhynchocyon_cirnei,PA44)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))))),(PA3,PA1)),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((PA5,((PA7,((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),((PA8,(PA21,(((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))),(Manis_pentadactyla,Metacheiromys_marshi)))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))),(PA6,((PA26,(PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))))),((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))))),((PA38,(PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(Orycteropus_afer,(PA43,(Rhynchocyon_cirnei,PA44)))))),((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)))),(PA3,PA1)),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,((((PA5,((PA6,(PA7,(((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(((Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))),(Leptictis_dakotensis,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),((PA9,((PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))),(PA8,(PA21,(Manis_pentadactyla,Metacheiromys_marshi))))))),((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA26,((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))))),((PA38,((PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(Orycteropus_afer,(PA43,(Rhynchocyon_cirnei,PA44))))),(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((((Solenodon_paradoxus,(Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea))),(Echinops_telfairi,Erinaceus_europaeus)),(Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))))),((PA5,((PA7,(((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),(PA8,(PA21,(((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA6,(PA26,((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA33,(((Cynocephalus_volans,Galeopterus_variegatus),(Ptilocercus_lowii,Tupaia_glis)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus))))))))),((PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),((Orycteropus_afer,PA43),(Rhynchocyon_cirnei,PA44)))),(PA38,(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((((Solenodon_paradoxus,(Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea))),(Echinops_telfairi,Erinaceus_europaeus)),(Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))))),((PA5,((PA7,(((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),(PA8,(PA21,(((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA6,(PA26,((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA33,(((Cynocephalus_volans,Galeopterus_variegatus),(Ptilocercus_lowii,Tupaia_glis)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus))))))))),((PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),((Orycteropus_afer,PA43),(Rhynchocyon_cirnei,PA44)))),(PA38,(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((Eomaia_scansoria,(((((PA6,(((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(((Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius)))),(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus)))),(Leptictis_dakotensis,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei))))),(PA7,((PA9,((PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))),(PA8,(PA21,(Manis_pentadactyla,Metacheiromys_marshi))))))),((PA26,(PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))))),((PA33,(Cynocephalus_volans,Galeopterus_variegatus)),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))),(PA5,((PA38,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA40,(PA43,(Orycteropus_afer,(Rhynchocyon_cirnei,PA44)))))),(Dasypus_novemcinctus,(PA39,(Hapalops_elongatus,Tamandua_tetradactyla)))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));

[“max preservation” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: strict consensus of 8 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.908000:0.000000,((Eomaia_scansoria:0.000000,Leptictis_dakotensis:0.000000,Maelestes_gobiensis:0.000000,PA21:0.000000,PA8:0.000000,PA7:0.000000,PA6:0.000000,PA5:0.000000,PA3:0.000000,PA1:0.000000,(Solenodon_paradoxus:0.000000,(Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.456000:0.000000)0.784000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.724000:0.000000)0.284000:0.000000,(Protungulatum_donnae:0.000000,(Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.344000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.244000:0.000000)0.100000:0.000000)0.136000:0.000000)0.192000:0.000000,(PA9:0.000000,(PA10:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.996000:0.000000)0.188000:0.000000)0.752000:0.000000)0.268000:0.000000)0.120000:0.000000)0.176000:0.000000)0.196000:0.000000)0.180000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.932000:0.000000)0.140000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.504000:0.000000)0.076000:0.000000)0.540000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.696000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.752000:0.000000)0.996000:0.000000)1.000000:0.000000)0.992000:0.000000)1.000000:0.000000)0.088000:0.000000,(Sinopa_rapax:0.000000,(Canis_lupus:0.000000,Felis_silvestris:0.000000)0.812000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)1.000000:0.000000,(PA38:0.000000,(PA40:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.240000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.108000:0.000000)0.116000:0.000000)0.040000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.460000:0.000000)0.016000:0.000000)0.052000:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000)0.868000:0.000000)0.644000:0.000000)0.024000:0.000000,(Dasypus_novemcinctus:0.000000,PA39:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.988000:0.000000)0.636000:0.000000)0.004000:0.000000,(PA26:0.000000,(PA27:0.000000,(PA28:0.000000,((Ictidomys_tridecemlineatus:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.896000:0.000000)0.284000:0.000000,(Rattus_norvegicus:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.328000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.656000:0.000000)1.000000:0.000000)0.316000:0.000000)0.292000:0.000000)0.160000:0.000000)0.116000:0.000000)0.812000:0.000000)1.000000:0.000000)0.100000:0.000000,(PA33:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.996000:0.000000)0.500000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.108000:0.000000)0.728000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)1.000000:0.000000)0.992000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.924000:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.608000:0.000000)0.888000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.872000:0.000000)0.860000:0.000000)1.000000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“max preservation” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: strict consensus of 8 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.908000:0.000000,((Eomaia_scansoria:0.000000,Leptictis_dakotensis:0.000000,Maelestes_gobiensis:0.000000,PA21:0.000000,PA8:0.000000,PA7:0.000000,PA6:0.000000,PA5:0.000000,PA3:0.000000,PA1:0.000000,(Solenodon_paradoxus:0.000000,(Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.456000:0.000000)0.840000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.724000:0.000000)0.596000:0.000000,(Protungulatum_donnae:0.000000,(Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.344000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.244000:0.000000)0.326667:0.000000)0.395000:0.000000)0.395200:0.000000,(PA9:0.000000,(PA10:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.996000:0.000000)0.592000:0.000000)0.856000:0.000000)0.714000:0.000000)0.615200:0.000000)0.578667:0.000000)0.549143:0.000000)0.510000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.932000:0.000000)0.555200:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.504000:0.000000)0.614667:0.000000)0.649538:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.696000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.752000:0.000000)0.998000:0.000000)1.000000:0.000000)0.998667:0.000000)1.000000:0.000000)0.508727:0.000000,(Sinopa_rapax:0.000000,(Canis_lupus:0.000000,Felis_silvestris:0.000000)0.812000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)1.000000:0.000000,(PA38:0.000000,(PA40:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.240000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.108000:0.000000)0.248000:0.000000)0.262000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.460000:0.000000)0.257333:0.000000)0.266857:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000)0.868000:0.000000)0.718667:0.000000)0.285000:0.000000,(Dasypus_novemcinctus:0.000000,PA39:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.988000:0.000000)0.754667:0.000000)0.232471:0.000000,(PA26:0.000000,(PA27:0.000000,(PA28:0.000000,((Ictidomys_tridecemlineatus:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.896000:0.000000)0.612000:0.000000,(Rattus_norvegicus:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.328000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.656000:0.000000)1.000000:0.000000)0.772000:0.000000)0.696000:0.000000)0.540667:0.000000)0.621714:0.000000)0.981200:0.000000)1.000000:0.000000)0.925000:0.000000,(PA33:0.000000,(Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.996000:0.000000)0.748000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.595111:0.000000)0.858087:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)1.000000:0.000000)0.994667:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.924000:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.608000:0.000000)0.930000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.872000:0.000000)0.930000:0.000000)1.000000:0.000000)1.000000:0.000000)-1.#IND00:0.000000);

[“max preservation” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: 4 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((PA3,(PA1,(PA5,((PA6,((PA7,((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(Leptictis_dakotensis,Rhynchocyon_cirnei)))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))),(PA38,(PA40,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Orycteropus_afer,(Manis_pentadactyla,Metacheiromys_marshi))))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((PA6,((PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus)))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))),(PA5,(((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(PA38,(PA40,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Orycteropus_afer,(Manis_pentadactyla,Metacheiromys_marshi)))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),(((((PA5,((PA6,((PA7,((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(Leptictis_dakotensis,Rhynchocyon_cirnei)))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))),(PA38,(PA40,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Orycteropus_afer,(Manis_pentadactyla,Metacheiromys_marshi))))))),(PA3,PA1)),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),(((((PA5,((PA6,((PA7,(Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus)))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))),(((((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(PA38,(PA40,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Orycteropus_afer,(Manis_pentadactyla,Metacheiromys_marshi)))))))),(PA3,PA1)),(Dromiciops_gliroides,(PA4,(Didelphis_virginiana,Pucadelphys_andinus)))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));

[“max preservation” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: strict consensus of 4 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.956000:0.000000,((((PA3:0.000000,PA1:0.000000,(PA7:0.000000,PA6:0.000000,PA5:0.000000,(Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.516000:0.000000)0.928000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.956000:0.000000)0.296000:0.000000)0.308000:0.000000,((((Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.316000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.448000:0.000000)0.080000:0.000000)0.328000:0.000000,(Protungulatum_donnae:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)1.000000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.812000:0.000000)1.000000:0.000000)0.068000:0.000000)0.044000:0.000000,(((Procavia_capensis:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)1.000000:0.000000)0.116000:0.000000)0.728000:0.000000)0.124000:0.000000)0.060000:0.000000)0.076000:0.000000)0.080000:0.000000)0.056000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.984000:0.000000)0.056000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.744000:0.000000)0.408000:0.000000)0.016000:0.000000,((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.016000:0.000000,(Moeritherium_lyonsi:0.000000,(Loxodonta_africana:0.000000,Trichechus_manatus:0.000000)0.140000:0.000000)0.408000:0.000000)0.048000:0.000000)0.280000:0.000000,((Rattus_norvegicus:0.000000,((Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.936000:0.000000,(Cocomys_lingchaensis:0.000000,Ictidomys_tridecemlineatus:0.000000)0.392000:0.000000)0.240000:0.000000)0.120000:0.000000,(Paramys_delicatus:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.940000:0.000000)1.000000:0.000000)0.412000:0.000000)0.284000:0.000000)0.028000:0.000000)1.000000:0.000000)0.060000:0.000000)0.196000:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.380000:0.000000)0.036000:0.000000,(PA26:0.000000,(((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.732000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.784000:0.000000)0.996000:0.000000)1.000000:0.000000)1.000000:0.000000)1.000000:0.000000)0.072000:0.000000,((Notharctus_tenebrosus:0.000000,(Lemur_catta:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.996000:0.000000)0.520000:0.000000)0.168000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.148000:0.000000)0.072000:0.000000)0.072000:0.000000,(PA38:0.000000,(PA40:0.000000,(((Dasypus_novemcinctus:0.000000,PA39:0.000000)0.240000:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.996000:0.000000)0.648000:0.000000,(Orycteropus_afer:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)1.000000:0.000000)0.276000:0.000000)0.236000:0.000000)0.060000:0.000000)0.316000:0.000000)0.436000:0.000000)0.416000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)1.000000:0.000000)1.000000:0.000000)0.576000:0.000000,(Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.864000:0.000000)0.472000:0.000000)0.948000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.936000:0.000000)0.876000:0.000000)1.000000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“max preservation” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: strict consensus of 4 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.956000:0.000000,((((PA3:0.000000,PA1:0.000000,(PA7:0.000000,PA6:0.000000,PA5:0.000000,(Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.516000:0.000000)0.950000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.956000:0.000000)0.664000:0.000000)0.628000:0.000000,((((Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.316000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.448000:0.000000)0.393333:0.000000)0.473000:0.000000,(Protungulatum_donnae:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)1.000000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.937333:0.000000)1.000000:0.000000)0.813600:0.000000)0.438400:0.000000,(((Procavia_capensis:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)1.000000:0.000000)0.558000:0.000000)0.829333:0.000000)0.654000:0.000000)0.544000:0.000000)0.481333:0.000000)0.434857:0.000000)0.422000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.984000:0.000000)0.498400:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.744000:0.000000)0.597000:0.000000)0.575385:0.000000,((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.016000:0.000000,(Moeritherium_lyonsi:0.000000,(Loxodonta_africana:0.000000,Trichechus_manatus:0.000000)0.140000:0.000000)0.584000:0.000000)0.536000:0.000000)0.564222:0.000000,((Rattus_norvegicus:0.000000,((Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.936000:0.000000,(Cocomys_lingchaensis:0.000000,Ictidomys_tridecemlineatus:0.000000)0.392000:0.000000)0.558667:0.000000)0.540000:0.000000,(Paramys_delicatus:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.940000:0.000000)1.000000:0.000000)0.804000:0.000000)0.728000:0.000000)0.591200:0.000000)1.000000:0.000000)0.514069:0.000000)0.497800:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.380000:0.000000)0.479619:0.000000,(PA26:0.000000,(((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.732000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.784000:0.000000)0.998000:0.000000)1.000000:0.000000)1.000000:0.000000)1.000000:0.000000)0.793778:0.000000,((Notharctus_tenebrosus:0.000000,(Lemur_catta:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.996000:0.000000)0.758000:0.000000)0.722667:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.716000:0.000000)0.509500:0.000000)0.515765:0.000000,(PA38:0.000000,(PA40:0.000000,(((Dasypus_novemcinctus:0.000000,PA39:0.000000)0.240000:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.996000:0.000000)0.765333:0.000000,(Orycteropus_afer:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)1.000000:0.000000)0.638000:0.000000)0.628000:0.000000)0.577714:0.000000)0.581500:0.000000)0.686250:0.000000)0.716286:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)1.000000:0.000000)1.000000:0.000000)0.819200:0.000000,(Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.864000:0.000000)0.720000:0.000000)0.965333:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.936000:0.000000)0.938000:0.000000)1.000000:0.000000)1.000000:0.000000)-1.#IND00:0.000000);

[“typical preservation" PAs corrected for the O'Leary et al. character ontology: 2 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Tachyglossus_aculeatus,(Ornithorhynchus_anatinus,PA2)),((((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),((Leptictis_dakotensis,(Zalambdalestes_lechei,(Ukhaatherium_nessovi,(Eomaia_scansoria,Maelestes_gobiensis)))),(Erinaceus_europaeus,((PA24,(Solenodon_paradoxus,PA23)),(Sorex_araneus,(Talpa_europaea,PA25)))))),((PA5,((PA7,(PA8,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((Sus_scrofa,(PA12,(PA13,(Hippopotamus_amphibius,(PA14,((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus)))))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,((Canis_lupus,(Felis_silvestris,PA22)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))))))),(PA6,((PA27,(PA28,((((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(PA26,(((Galeopterus_variegatus,(Cynocephalus_volans,PA34)),(Ptilocercus_lowii,(Tupaia_glis,PA32))),(PA33,((PA35,(Tarsius_syrichta,(PA36,(PA37,(Homo_sapiens,Saimiri_sciureus))))),(Lemur_catta,Notharctus_tenebrosus))))))))),(PA38,((PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),((Orycteropus_afer,PA43),(Rhynchocyon_cirnei,PA44)))),(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA1,(PA3,((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4)))))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,(((Tachyglossus_aculeatus,(Ornithorhynchus_anatinus,(PA2,PA1))),((((Echinops_telfairi,(Amblysomus_hottentotus,PA45)),((Leptictis_dakotensis,(Zalambdalestes_lechei,(Ukhaatherium_nessovi,(Eomaia_scansoria,Maelestes_gobiensis)))),(Erinaceus_europaeus,((Solenodon_paradoxus,PA23),(PA24,(PA25,(Sorex_araneus,Talpa_europaea))))))),((PA5,(PA6,((PA7,(PA8,((((((Apheliscus_insidiosus,Protungulatum_donnae),((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Rodhocetus_balochistanensis,(Archaeotherium_mortoni,Mesonyx_obtusidens))),((PA10,((Sus_scrofa,(PA12,(PA13,(((Artiocetus_clavis,Basilosaurus_cetoides),(PA15,(Caperea_marginata,Tursiops_truncatus))),(Hippopotamus_amphibius,PA14))))),(PA11,(Bos_taurus,Lama_glama)))),(Equus_caballus,Mesohippus_bairdi))),(PA9,(((Icaronycteris_index,Onychonycteris_finneyi),(((Myotis_lucifugus,PA17),(Pteronotus_parnellii,PA18)),((Nycteris_thebaica,PA19),(Rhinopoma_hardwickii,Saccopteryx_bilineata)))),(PA16,(Pteropus_giganteus,PA20))))),(PA21,((PA22,(Canis_lupus,Felis_silvestris)),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))))))),((PA27,(PA28,((((Castor_canadensis,Cavia_porcellus),(Ictidomys_tridecemlineatus,PA31)),(PA29,(Rattus_norvegicus,PA30))),((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))),(PA26,(((Cynocephalus_volans,(Galeopterus_variegatus,PA34)),(Ptilocercus_lowii,(Tupaia_glis,PA32))),(PA33,(PA35,((PA36,(Tarsius_syrichta,(Homo_sapiens,(Saimiri_sciureus,PA37)))),(Lemur_catta,Notharctus_tenebrosus)))))))))),(PA38,(((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA40,((Orycteropus_afer,PA43),(Rhynchocyon_cirnei,PA44)))),(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA3,(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden)))));

[“typical preservation” PAs corrected for the O'Leary et al. character ontology: strict consensus of 2 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000,PA3:0.000000,PA2:0.000000,PA1:0.000000,(((Echinops_telfairi:0.000000,(Amblysomus_hottentotus:0.000000,PA45:0.000000)0.048000:0.000000)0.524000:0.000000,((Leptictis_dakotensis:0.000000,(Zalambdalestes_lechei:0.000000,(Ukhaatherium_nessovi:0.000000,(Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000)0.432000:0.000000)0.120000:0.000000)0.596000:0.000000)0.040000:0.000000,(Erinaceus_europaeus:0.000000,(PA24:0.000000,(Solenodon_paradoxus:0.000000,PA23:0.000000)0.020000:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000,PA25:0.000000)0.852000:0.000000)0.024000:0.000000)0.152000:0.000000)0.008000:0.000000)0.000000:0.000000,((PA5:0.000000,(PA6:0.000000,(PA7:0.000000,(PA8:0.000000,(((((Apheliscus_insidiosus:0.000000,Protungulatum_donnae:0.000000)0.036000:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.316000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.228000:0.000000)0.060000:0.000000)0.208000:0.000000,(Rodhocetus_balochistanensis:0.000000,(Archaeotherium_mortoni:0.000000,Mesonyx_obtusidens:0.000000)0.060000:0.000000)0.032000:0.000000)0.000000:0.000000,((PA10:0.000000,(PA11:0.000000,(Sus_scrofa:0.000000,PA12:0.000000,(PA13:0.000000,(Hippopotamus_amphibius:0.000000,PA14:0.000000,((Artiocetus_clavis:0.000000,Basilosaurus_cetoides:0.000000)0.180000:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000,PA15:0.000000)0.992000:0.000000)0.200000:0.000000)0.176000:0.000000)0.016000:0.000000)0.012000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.044000:0.000000)0.012000:0.000000)0.000000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.292000:0.000000)0.028000:0.000000)0.000000:0.000000,(PA9:0.000000,(((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,(((Myotis_lucifugus:0.000000,PA17:0.000000)0.348000:0.000000,(Pteronotus_parnellii:0.000000,PA18:0.000000)0.208000:0.000000)0.568000:0.000000,((Nycteris_thebaica:0.000000,PA19:0.000000)0.196000:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.632000:0.000000)0.900000:0.000000)0.752000:0.000000)0.408000:0.000000,(PA16:0.000000,(Pteropus_giganteus:0.000000,PA20:0.000000)0.396000:0.000000)0.212000:0.000000)1.000000:0.000000)0.428000:0.000000)0.000000:0.000000,(PA21:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000,PA22:0.000000)0.764000:0.000000,(Sinopa_rapax:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.668000:0.000000)0.720000:0.000000)0.184000:0.000000)0.000000:0.000000)0.000000:0.000000,(PA26:0.000000,(PA27:0.000000,(PA28:0.000000,((PA29:0.000000,(Ictidomys_tridecemlineatus:0.000000,PA31:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.264000:0.000000)0.036000:0.000000,(Rattus_norvegicus:0.000000,PA30:0.000000)0.156000:0.000000)0.068000:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.456000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.736000:0.000000)0.996000:0.000000)0.360000:0.000000)0.396000:0.000000)0.160000:0.000000)0.252000:0.000000)1.000000:0.000000)0.016000:0.000000,(PA33:0.000000,((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000,PA34:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,(Tupaia_glis:0.000000,PA32:0.000000)0.016000:0.000000)1.000000:0.000000)0.012000:0.000000,(PA35:0.000000,(Tarsius_syrichta:0.000000,PA36:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000,PA37:0.000000)1.000000:0.000000)0.400000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000)0.208000:0.000000)1.000000:0.000000)0.016000:0.000000)0.216000:0.000000)0.000000:0.000000)0.000000:0.000000,(PA38:0.000000,((PA40:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.176000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.100000:0.000000)0.052000:0.000000)0.032000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.256000:0.000000)0.004000:0.000000)0.044000:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000)0.048000:0.000000)0.028000:0.000000)0.004000:0.000000,(((Dasypus_novemcinctus:0.000000,PA39:0.000000)0.128000:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.980000:0.000000)0.476000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.972000:0.000000)0.304000:0.000000)0.000000:0.000000)0.004000:0.000000)0.000000:0.000000)0.060000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.988000:0.000000)0.992000:0.000000)0.472000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.964000:0.000000)1.000000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“typical preservation” PAs corrected for the O'Leary et al. character ontology: strict consensus of 2 MPTs - nodal values = "transfer bootstrap expectation" values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000,PA3:0.000000,PA2:0.000000,PA1:0.000000,(((Echinops_telfairi:0.000000,(Amblysomus_hottentotus:0.000000,PA45:0.000000)0.048000:0.000000)0.550000:0.000000,((Leptictis_dakotensis:0.000000,(Zalambdalestes_lechei:0.000000,(Ukhaatherium_nessovi:0.000000,(Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000)0.432000:0.000000)0.364000:0.000000)0.688000:0.000000)0.534000:0.000000,(Erinaceus_europaeus:0.000000,(PA24:0.000000,(Solenodon_paradoxus:0.000000,PA23:0.000000)0.020000:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000,PA25:0.000000)0.858000:0.000000)0.449600:0.000000)0.447333:0.000000)0.308364:0.000000)0.246286:0.000000,((PA5:0.000000,(PA6:0.000000,(PA7:0.000000,(PA8:0.000000,(((((Apheliscus_insidiosus:0.000000,Protungulatum_donnae:0.000000)0.036000:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.316000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.228000:0.000000)0.261333:0.000000)0.336800:0.000000,(Rodhocetus_balochistanensis:0.000000,(Archaeotherium_mortoni:0.000000,Mesonyx_obtusidens:0.000000)0.060000:0.000000)0.070000:0.000000)0.216500:0.000000,((PA10:0.000000,(PA11:0.000000,(Sus_scrofa:0.000000,PA12:0.000000,(PA13:0.000000,(Hippopotamus_amphibius:0.000000,PA14:0.000000,((Artiocetus_clavis:0.000000,Basilosaurus_cetoides:0.000000)0.180000:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000,PA15:0.000000)0.992000:0.000000)0.640000:0.000000)0.516667:0.000000)0.445143:0.000000)0.351111:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.044000:0.000000)0.407333:0.000000)0.450154:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.292000:0.000000)0.524533:0.000000)0.462500:0.000000,(PA9:0.000000,(((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,(((Myotis_lucifugus:0.000000,PA17:0.000000)0.348000:0.000000,(Pteronotus_parnellii:0.000000,PA18:0.000000)0.208000:0.000000)0.592000:0.000000,((Nycteris_thebaica:0.000000,PA19:0.000000)0.196000:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.632000:0.000000)0.917333:0.000000)0.860000:0.000000)0.844000:0.000000,(PA16:0.000000,(Pteropus_giganteus:0.000000,PA20:0.000000)0.396000:0.000000)0.312000:0.000000)1.000000:0.000000)0.956000:0.000000)0.401684:0.000000,(PA21:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000,PA22:0.000000)0.766000:0.000000,(Sinopa_rapax:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.834000:0.000000)0.848800:0.000000)0.746667:0.000000)0.336609:0.000000)0.329702:0.000000,(PA26:0.000000,(PA27:0.000000,(PA28:0.000000,((PA29:0.000000,(Ictidomys_tridecemlineatus:0.000000,PA31:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.264000:0.000000)0.125333:0.000000,(Rattus_norvegicus:0.000000,PA30:0.000000)0.156000:0.000000)0.184667:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.456000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.736000:0.000000)0.996000:0.000000)0.784000:0.000000)0.763000:0.000000)0.592000:0.000000)0.942462:0.000000)1.000000:0.000000)0.934400:0.000000,(PA33:0.000000,((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000,PA34:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,(Tupaia_glis:0.000000,PA32:0.000000)0.016000:0.000000)1.000000:0.000000)0.407200:0.000000,(PA35:0.000000,(Tarsius_syrichta:0.000000,PA36:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000,PA37:0.000000)1.000000:0.000000)0.750000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000)0.208000:0.000000)1.000000:0.000000)0.700286:0.000000)0.594065:0.000000)0.156898:0.000000)0.160000:0.000000,(PA38:0.000000,((PA40:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.176000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.100000:0.000000)0.162000:0.000000)0.196000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.256000:0.000000)0.184000:0.000000)0.186857:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000)0.048000:0.000000)0.036000:0.000000)0.114333:0.000000,(((Dasypus_novemcinctus:0.000000,PA39:0.000000)0.128000:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.980000:0.000000)0.648000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.972000:0.000000)0.604000:0.000000)0.192000:0.000000)0.183368:0.000000)0.253429:0.000000)0.410154:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.988000:0.000000)0.992000:0.000000)0.736000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.964000:0.000000)1.000000:0.000000)1.000000:0.000000));

[“typical preservation” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: 4 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),((PA6,((PA7,(PA8,((PA21,((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))),(PA26,((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA33,(((Cynocephalus_volans,Galeopterus_variegatus),(Ptilocercus_lowii,Tupaia_glis)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus)))))))),(PA5,(PA38,(((PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(Orycteropus_afer,(PA43,(Rhynchocyon_cirnei,PA44)))),(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(((PA7,((PA21,((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))))),(PA8,((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))),(PA6,((PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus))))))))),(PA26,(PA33,(((Cynocephalus_volans,Galeopterus_variegatus),(Ptilocercus_lowii,Tupaia_glis)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus)))))))),(PA5,(PA38,((PA40,((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA43,(Orycteropus_afer,(Rhynchocyon_cirnei,PA44))))),(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi))))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),((PA5,((PA7,((PA8,(PA21,((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus)))))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))))),(PA6,((PA26,(PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))))),(PA33,(((Cynocephalus_volans,Galeopterus_variegatus),(Ptilocercus_lowii,Tupaia_glis)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus)))))))),(PA38,(((PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42))),(PA40,((Orycteropus_afer,PA43),(Rhynchocyon_cirnei,PA44)))),(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),((PA5,((PA7,(PA8,((PA21,((Leptictis_dakotensis,(Protungulatum_donnae,(Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))))),((Canis_lupus,Felis_silvestris),(Sinopa_rapax,(Vulpavus_ovatus,Vulpavus_profectus))))),((PA9,(PA10,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi)))),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata))))))))),(PA6,((PA26,(PA27,(PA28,((Ictidomys_tridecemlineatus,(Castor_canadensis,Cavia_porcellus)),(Rattus_norvegicus,((Cocomys_lingchaensis,Paramys_delicatus),(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))))),(PA33,(((Cynocephalus_volans,Galeopterus_variegatus),(Ptilocercus_lowii,Tupaia_glis)),((Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)),(Lemur_catta,Notharctus_tenebrosus)))))))),(PA38,(((PA40,(PA41,(((Carodnia_vieirai,Thomashuxleya_externa_comb),(Loxodonta_africana,(Moeritherium_lyonsi,Trichechus_manatus))),(Procavia_capensis,PA42)))),(PA43,(Orycteropus_afer,(Rhynchocyon_cirnei,PA44)))),(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Manis_pentadactyla,Metacheiromys_marshi)))))),(PA3,PA1)),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));

[“typical preservation” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: strict consensus of 4 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.952000:0.000000,((((((Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.476000:0.000000)0.808000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.768000:0.000000)0.200000:0.000000)0.300000:0.000000,(PA5:0.000000,(PA6:0.000000,(PA7:0.000000,(PA8:0.000000,(PA21:0.000000,((Leptictis_dakotensis:0.000000,(Protungulatum_donnae:0.000000,(Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.316000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.260000:0.000000)0.068000:0.000000)0.128000:0.000000)0.144000:0.000000)0.000000:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)0.848000:0.000000,(Sinopa_rapax:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.064000:0.000000)0.976000:0.000000)0.000000:0.000000)0.000000:0.000000,((PA9:0.000000,(PA10:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.992000:0.000000)0.108000:0.000000)0.644000:0.000000)0.116000:0.000000)0.024000:0.000000)0.036000:0.000000)0.044000:0.000000)0.016000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.928000:0.000000)0.016000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.484000:0.000000)0.004000:0.000000)0.428000:0.000000)0.004000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.716000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.700000:0.000000)0.992000:0.000000)1.000000:0.000000)0.992000:0.000000)1.000000:0.000000)0.004000:0.000000)0.000000:0.000000)0.000000:0.000000,(PA26:0.000000,(PA27:0.000000,(PA28:0.000000,((Ictidomys_tridecemlineatus:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.848000:0.000000)0.232000:0.000000,(Rattus_norvegicus:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.300000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.652000:0.000000)1.000000:0.000000)0.292000:0.000000)0.320000:0.000000)0.088000:0.000000)0.044000:0.000000)0.448000:0.000000)1.000000:0.000000)0.048000:0.000000,(PA33:0.000000,(((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.000000:0.000000,((Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.976000:0.000000)0.400000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000)0.212000:0.000000)1.000000:0.000000)0.000000:0.000000)0.060000:0.000000)0.136000:0.000000)0.000000:0.000000,(PA38:0.000000,((PA40:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.120000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.080000:0.000000)0.112000:0.000000)0.024000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.308000:0.000000)0.000000:0.000000)0.012000:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000)0.764000:0.000000)0.328000:0.000000)0.000000:0.000000,(((Dasypus_novemcinctus:0.000000,PA39:0.000000)0.128000:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.992000:0.000000)0.544000:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.948000:0.000000)0.296000:0.000000)0.000000:0.000000)0.000000:0.000000)0.000000:0.000000)0.184000:0.000000,(PA3:0.000000,PA1:0.000000)0.048000:0.000000)0.220000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.948000:0.000000)0.984000:0.000000)0.308000:0.000000,(Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.652000:0.000000)0.256000:0.000000)0.936000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.796000:0.000000)0.904000:0.000000)1.000000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“typical preservation” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: strict consensus of 4 MPTs - nodal values = "transfer bootstrap expectation" bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.952000:0.000000,((((((Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.476000:0.000000)0.862000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.768000:0.000000)0.609000:0.000000)0.605600:0.000000,(PA5:0.000000,(PA6:0.000000,(PA7:0.000000,(PA8:0.000000,(PA21:0.000000,((Leptictis_dakotensis:0.000000,(Protungulatum_donnae:0.000000,(Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.316000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.260000:0.000000)0.282667:0.000000)0.336000:0.000000)0.324000:0.000000)0.270000:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)0.848000:0.000000,(Sinopa_rapax:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.532000:0.000000)0.982000:0.000000)0.382182:0.000000)0.350667:0.000000,((PA9:0.000000,(PA10:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.992000:0.000000)0.552000:0.000000)0.782667:0.000000)0.618000:0.000000)0.504000:0.000000)0.434667:0.000000)0.385714:0.000000)0.352000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.928000:0.000000)0.430000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.484000:0.000000)0.515000:0.000000)0.560308:0.000000)0.520571:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.716000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.700000:0.000000)0.994000:0.000000)1.000000:0.000000)0.998667:0.000000)1.000000:0.000000)0.448182:0.000000)0.278556:0.000000)0.271351:0.000000,(PA26:0.000000,(PA27:0.000000,(PA28:0.000000,((Ictidomys_tridecemlineatus:0.000000,(Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.848000:0.000000)0.578000:0.000000,(Rattus_norvegicus:0.000000,((Cocomys_lingchaensis:0.000000,Paramys_delicatus:0.000000)0.300000:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.652000:0.000000)1.000000:0.000000)0.764000:0.000000)0.709000:0.000000)0.522667:0.000000)0.553143:0.000000)0.944800:0.000000)1.000000:0.000000)0.920667:0.000000,(PA33:0.000000,(((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.333333:0.000000,((Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.976000:0.000000)0.688000:0.000000,(Lemur_catta:0.000000,Notharctus_tenebrosus:0.000000)0.212000:0.000000)1.000000:0.000000)0.559500:0.000000)0.565778:0.000000)0.549217:0.000000)0.206977:0.000000,(PA38:0.000000,((PA40:0.000000,(PA41:0.000000,(((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.120000:0.000000,(Loxodonta_africana:0.000000,(Moeritherium_lyonsi:0.000000,Trichechus_manatus:0.000000)0.080000:0.000000)0.204000:0.000000)0.202000:0.000000,(Procavia_capensis:0.000000,PA42:0.000000)0.308000:0.000000)0.176000:0.000000)0.168000:0.000000,(Orycteropus_afer:0.000000,PA43:0.000000,(Rhynchocyon_cirnei:0.000000,PA44:0.000000)0.764000:0.000000)0.477333:0.000000)0.170667:0.000000,(((Dasypus_novemcinctus:0.000000,PA39:0.000000)0.128000:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.992000:0.000000)0.694667:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.948000:0.000000)0.626400:0.000000)0.196667:0.000000)0.187158:0.000000)0.401273:0.000000)0.549250:0.000000,(PA3:0.000000,PA1:0.000000)0.048000:0.000000)0.594000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.948000:0.000000)0.988000:0.000000)0.718800:0.000000,(Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.652000:0.000000)0.518667:0.000000)0.960667:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.796000:0.000000)0.952000:0.000000)1.000000:0.000000)1.000000:0.000000)-1.#IND00:0.000000);

[“typical preservation” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: 2 MPTs]
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((((PA6,((PA7,(((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis)))))),(PA5,(PA38,(PA40,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Orycteropus_afer,(Manis_pentadactyla,Metacheiromys_marshi))))))),(PA3,PA1)),(PA4,(Dromiciops_gliroides,(Didelphis_virginiana,Pucadelphys_andinus)))),(Eomaia_scansoria,(Maelestes_gobiensis,(Ukhaatherium_nessovi,Zalambdalestes_lechei)))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));
(Morganucodon_oehleri,(Morganucodon_watsoni,(Haldanodon_exspectatus,((Ornithorhynchus_anatinus,Tachyglossus_aculeatus),((((PA1,(PA3,((PA5,(PA6,((PA7,(((Solenodon_paradoxus,((Amblysomus_hottentotus,(Sorex_araneus,Talpa_europaea)),(Echinops_telfairi,Erinaceus_europaeus))),(Leptictis_dakotensis,Rhynchocyon_cirnei)),(((Apheliscus_insidiosus,((Didolodus_multicuspis,Protolipterna_ellipsodontoides),(Hyopsodus_paulus,Phenacodus_intermedius))),(Protungulatum_donnae,(Sinopa_rapax,((Canis_lupus,Felis_silvestris),(Vulpavus_ovatus,Vulpavus_profectus))))),(((Procavia_capensis,(((Sus_scrofa,(Hippopotamus_amphibius,(Archaeotherium_mortoni,(Mesonyx_obtusidens,(Rodhocetus_balochistanensis,(Basilosaurus_cetoides,(Artiocetus_clavis,(Caperea_marginata,Tursiops_truncatus)))))))),(Bos_taurus,Lama_glama)),(Equus_caballus,Mesohippus_bairdi))),((Carodnia_vieirai,Thomashuxleya_externa_comb),(Moeritherium_lyonsi,(Loxodonta_africana,Trichechus_manatus)))),((Rattus_norvegicus,((Castor_canadensis,Cavia_porcellus),(Cocomys_lingchaensis,Ictidomys_tridecemlineatus))),(Paramys_delicatus,(Tribosphenomys_minutus,(Rhombomylus_turpanensis,(Gomphos_elkema,(Dawsonolagus_antiquus,Oryctolagus_cuniculus)))))))))),(PA26,(((Cynocephalus_volans,Galeopterus_variegatus),(Pteropus_giganteus,((Icaronycteris_index,Onychonycteris_finneyi),((Myotis_lucifugus,Pteronotus_parnellii),(Nycteris_thebaica,(Rhinopoma_hardwickii,Saccopteryx_bilineata)))))),((Notharctus_tenebrosus,(Lemur_catta,(Tarsius_syrichta,(Homo_sapiens,Saimiri_sciureus)))),(Ptilocercus_lowii,Tupaia_glis))))))),(PA38,(PA40,(((Dasypus_novemcinctus,PA39),(Hapalops_elongatus,Tamandua_tetradactyla)),(Orycteropus_afer,(Manis_pentadactyla,Metacheiromys_marshi)))))))),((Didelphis_virginiana,Pucadelphys_andinus),(Dromiciops_gliroides,PA4))),((Eomaia_scansoria,Maelestes_gobiensis),(Ukhaatherium_nessovi,Zalambdalestes_lechei))),(Henkelotherium_guimarotae,Zhangheotherium_quinquecuspiden))))));

[“typical preservation” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: strict consensus of 2 MPTs - nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.968000:0.000000,((((PA3:0.000000,PA1:0.000000,(PA5:0.000000,(PA6:0.000000,((PA7:0.000000,(((Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.480000:0.000000)0.832000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.824000:0.000000)0.248000:0.000000)0.272000:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.396000:0.000000)0.052000:0.000000,(((Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.276000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.348000:0.000000)0.104000:0.000000)0.256000:0.000000,(Protungulatum_donnae:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)0.992000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.644000:0.000000)0.996000:0.000000)0.064000:0.000000)0.036000:0.000000,(((Procavia_capensis:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.992000:0.000000)0.088000:0.000000)0.560000:0.000000)0.072000:0.000000)0.024000:0.000000)0.056000:0.000000)0.032000:0.000000)0.016000:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.984000:0.000000)0.020000:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.596000:0.000000)0.264000:0.000000)0.008000:0.000000,((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.024000:0.000000,(Moeritherium_lyonsi:0.000000,(Loxodonta_africana:0.000000,Trichechus_manatus:0.000000)0.144000:0.000000)0.324000:0.000000)0.020000:0.000000)0.172000:0.000000,((Rattus_norvegicus:0.000000,((Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.844000:0.000000,(Cocomys_lingchaensis:0.000000,Ictidomys_tridecemlineatus:0.000000)0.384000:0.000000)0.220000:0.000000)0.100000:0.000000,(Paramys_delicatus:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.852000:0.000000)1.000000:0.000000)0.444000:0.000000)0.260000:0.000000)0.052000:0.000000)1.000000:0.000000)0.080000:0.000000)0.160000:0.000000)0.004000:0.000000)0.004000:0.000000,(PA26:0.000000,(((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.684000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.744000:0.000000)0.980000:0.000000)1.000000:0.000000)0.992000:0.000000)1.000000:0.000000)0.056000:0.000000,((Notharctus_tenebrosus:0.000000,(Lemur_catta:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.988000:0.000000)0.456000:0.000000)0.160000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.136000:0.000000)0.032000:0.000000)0.060000:0.000000)0.004000:0.000000)0.044000:0.000000,(PA38:0.000000,(PA40:0.000000,(((Dasypus_novemcinctus:0.000000,PA39:0.000000)0.200000:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.972000:0.000000)0.444000:0.000000,(Orycteropus_afer:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.976000:0.000000)0.108000:0.000000)0.220000:0.000000)0.084000:0.000000)0.432000:0.000000)0.216000:0.000000)0.280000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.944000:0.000000)0.984000:0.000000)0.508000:0.000000,(Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.740000:0.000000)0.388000:0.000000)0.960000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.852000:0.000000)0.880000:0.000000)0.996000:0.000000)1.000000:0.000000)2.000000:0.000000);

[“typical preservation” PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: strict consensus of 2 MPTs - nodal values = "transfer bootstrap expectation" bootstrap values]
(Morganucodon_oehleri:0.000000,(Morganucodon_watsoni:0.000000,(Haldanodon_exspectatus:0.000000,((Ornithorhynchus_anatinus:0.000000,Tachyglossus_aculeatus:0.000000)0.968000:0.000000,((((PA3:0.000000,PA1:0.000000,(PA5:0.000000,(PA6:0.000000,((PA7:0.000000,(((Solenodon_paradoxus:0.000000,((Amblysomus_hottentotus:0.000000,(Sorex_araneus:0.000000,Talpa_europaea:0.000000)0.480000:0.000000)0.882000:0.000000,(Echinops_telfairi:0.000000,Erinaceus_europaeus:0.000000)0.824000:0.000000)0.635000:0.000000)0.612000:0.000000,(Leptictis_dakotensis:0.000000,Rhynchocyon_cirnei:0.000000)0.396000:0.000000)0.466286:0.000000,(((Apheliscus_insidiosus:0.000000,((Didolodus_multicuspis:0.000000,Protolipterna_ellipsodontoides:0.000000)0.276000:0.000000,(Hyopsodus_paulus:0.000000,Phenacodus_intermedius:0.000000)0.348000:0.000000)0.364000:0.000000)0.421000:0.000000,(Protungulatum_donnae:0.000000,(Sinopa_rapax:0.000000,((Canis_lupus:0.000000,Felis_silvestris:0.000000)0.992000:0.000000,(Vulpavus_ovatus:0.000000,Vulpavus_profectus:0.000000)1.000000:0.000000)0.881333:0.000000)0.999000:0.000000)0.812000:0.000000)0.430000:0.000000,(((Procavia_capensis:0.000000,(((Sus_scrofa:0.000000,(Hippopotamus_amphibius:0.000000,(Archaeotherium_mortoni:0.000000,(Mesonyx_obtusidens:0.000000,(Rodhocetus_balochistanensis:0.000000,(Basilosaurus_cetoides:0.000000,(Artiocetus_clavis:0.000000,(Caperea_marginata:0.000000,Tursiops_truncatus:0.000000)0.992000:0.000000)0.542000:0.000000)0.729333:0.000000)0.567000:0.000000)0.470400:0.000000)0.417333:0.000000)0.372571:0.000000)0.356500:0.000000,(Bos_taurus:0.000000,Lama_glama:0.000000)0.984000:0.000000)0.399200:0.000000,(Equus_caballus:0.000000,Mesohippus_bairdi:0.000000)0.596000:0.000000)0.470333:0.000000)0.451077:0.000000,((Carodnia_vieirai:0.000000,Thomashuxleya_externa_comb:0.000000)0.024000:0.000000,(Moeritherium_lyonsi:0.000000,(Loxodonta_africana:0.000000,Trichechus_manatus:0.000000)0.144000:0.000000)0.528000:0.000000)0.496000:0.000000)0.440667:0.000000,((Rattus_norvegicus:0.000000,((Castor_canadensis:0.000000,Cavia_porcellus:0.000000)0.844000:0.000000,(Cocomys_lingchaensis:0.000000,Ictidomys_tridecemlineatus:0.000000)0.384000:0.000000)0.530667:0.000000)0.514000:0.000000,(Paramys_delicatus:0.000000,(Tribosphenomys_minutus:0.000000,(Rhombomylus_turpanensis:0.000000,(Gomphos_elkema:0.000000,(Dawsonolagus_antiquus:0.000000,Oryctolagus_cuniculus:0.000000)0.852000:0.000000)1.000000:0.000000)0.814667:0.000000)0.719000:0.000000)0.588800:0.000000)1.000000:0.000000)0.499172:0.000000)0.467100:0.000000)0.407478:0.000000)0.397422:0.000000,(PA26:0.000000,(((Cynocephalus_volans:0.000000,Galeopterus_variegatus:0.000000)1.000000:0.000000,(Pteropus_giganteus:0.000000,((Icaronycteris_index:0.000000,Onychonycteris_finneyi:0.000000)1.000000:0.000000,((Myotis_lucifugus:0.000000,Pteronotus_parnellii:0.000000)0.684000:0.000000,(Nycteris_thebaica:0.000000,(Rhinopoma_hardwickii:0.000000,Saccopteryx_bilineata:0.000000)0.744000:0.000000)0.986000:0.000000)1.000000:0.000000)0.998667:0.000000)1.000000:0.000000)0.790222:0.000000,((Notharctus_tenebrosus:0.000000,(Lemur_catta:0.000000,(Tarsius_syrichta:0.000000,(Homo_sapiens:0.000000,Saimiri_sciureus:0.000000)0.988000:0.000000)0.722000:0.000000)0.720000:0.000000)1.000000:0.000000,(Ptilocercus_lowii:0.000000,Tupaia_glis:0.000000)1.000000:0.000000)0.712000:0.000000)0.497250:0.000000)0.509176:0.000000)0.397333:0.000000)0.412462:0.000000,(PA38:0.000000,(PA40:0.000000,(((Dasypus_novemcinctus:0.000000,PA39:0.000000)0.200000:0.000000,(Hapalops_elongatus:0.000000,Tamandua_tetradactyla:0.000000)0.972000:0.000000)0.625333:0.000000,(Orycteropus_afer:0.000000,(Manis_pentadactyla:0.000000,Metacheiromys_marshi:0.000000)0.976000:0.000000)0.542000:0.000000)0.634667:0.000000)0.605714:0.000000)0.631000:0.000000)0.610750:0.000000)0.662000:0.000000,(Dromiciops_gliroides:0.000000,PA4:0.000000,(Didelphis_virginiana:0.000000,Pucadelphys_andinus:0.000000)0.944000:0.000000)0.989333:0.000000)0.800800:0.000000,(Eomaia_scansoria:0.000000,Maelestes_gobiensis:0.000000,(Ukhaatherium_nessovi:0.000000,Zalambdalestes_lechei:0.000000)0.740000:0.000000)0.626667:0.000000)0.976000:0.000000,(Henkelotherium_guimarotae:0.000000,Zhangheotherium_quinquecuspiden:0.000000)0.852000:0.000000)0.940000:0.000000)0.998000:0.000000)1.000000:0.000000)-1.#IND00:0.000000);



[Maximum likelihood analyses]

[no PAs: nodal values = standard bootstrap values]
(Morganucodon_oehleri:0.007443,(Morganucodon_watsoni:0.008777,(Haldanodon_exspectatus:0.099164,(((Eomaia_scansoria:0.06223,((Dromiciops_gliroides:0.084822,(Pucadelphys_andinus:0.114211,Didelphis_virginiana:0.053163)'0.936111':0.054776)'1.000000':0.073516,((Maelestes_gobiensis:0.103069,(Zalambdalestes_lechei:0.124749,Ukhaatherium_nessovi:0.103981)'0.983333':0.055483)'0.622222':0.043684,((((Sinopa_rapax:0.089923,((Vulpavus_ovatus:0.017136,Vulpavus_profectus:0.02736)'1.000000':0.054969,(Felis_silvestris:0.09397,Canis_lupus:0.068977)'1.000000':0.061354)'0.886111':0.030009)'1.000000':0.077075,(Protungulatum_donnae:0.135793,((((Cocomys_lingchaensis:0.074623,Paramys_delicatus:0.05218)'0.355556':0.018293,(Rattus_norvegicus:0.151467,(Ictidomys_tridecemlineatus:0.063015,(Cavia_porcellus:0.090784,Castor_canadensis:0.086457)'0.947222':0.049951)'0.597222':0.0461)'0.477778':0.027639)'0.744444':0.033041,(Tribosphenomys_minutus:0.104868,(Rhombomylus_turpanensis:0.074478,(Gomphos_elkema:0.077086,(Dawsonolagus_antiquus:0.025474,Oryctolagus_cuniculus:0.098581)'0.975000':0.035809)'1.000000':0.072567)'0.616667':0.024996)'0.683333':0.033814)'1.000000':0.095354,((Apheliscus_insidiosus:0.114763,((Didolodus_multicuspis:0.070325,Protolipterna_ellipsodontoides:0.057358)'0.600000':0.033642,(Hyopsodus_paulus:0.100402,Phenacodus_intermedius:0.092737)'0.650000':0.026091)'0.338889':0.015702)'0.736111':0.026464,(((Mesonyx_obtusidens:0.155465,(Archaeotherium_mortoni:0.125507,(Rodhocetus_balochistanensis:0.104617,(Artiocetus_clavis:0.118052,(Basilosaurus_cetoides:0.141697,(Tursiops_truncatus:0.165577,Caperea_marginata:0.167985)'1.000000':0.113237)'0.583333':0.048819)'0.922222':0.051828)'0.591667':0.049302)'0.063889':0.026003)'0.347222':0.046651,((Equus_caballus:0.1459,Mesohippus_bairdi:0.151273)'0.866667':0.059552,((Hippopotamus_amphibius:0.19223,Sus_scrofa:0.160792)'0.225000':0.037404,(Lama_glama:0.121042,Bos_taurus:0.126859)'0.991667':0.060059)'0.119444':0.032286)'0.230556':0.044061)'0.808333':0.049429,((Procavia_capensis:0.194301,Thomashuxleya_externa_comb:0.13528)'0.302778':0.029765,(Carodnia_vieirai:0.087573,(Moeritherium_lyonsi:0.136601,(Trichechus_manatus:0.202685,Loxodonta_africana:0.168062)'0.505556':0.037237)'0.658333':0.042398)'0.963889':0.057215)'0.325000':0.04498)'0.791667':0.041203)'0.444444':0.035076)'0.438889':0.022444)'0.402778':0.019555)'0.430556':0.032656,((Orycteropus_afer:0.239321,((Metacheiromys_marshi:0.114432,Manis_pentadactyla:0.191407)'0.955556':0.059411,(Dasypus_novemcinctus:0.188778,(Hapalops_elongatus:0.157304,Tamandua_tetradactyla:0.142909)'1.000000':0.075132)'0.963889':0.055729)'0.936111':0.043583)'0.958333':0.06214,((Pteropus_giganteus:0.158564,((Onychonycteris_finneyi:0.031028,Icaronycteris_index:0.026973)'0.997222':0.060668,((Nycteris_thebaica:0.091164,(Rhinopoma_hardwickii:0.052008,Saccopteryx_bilineata:0.0521)'0.830556':0.025888)'0.986111':0.032396,(Myotis_lucifugus:0.061787,Pteronotus_parnellii:0.0761)'0.766667':0.026288)'1.000000':0.05286)'0.994444':0.05458)'1.000000':0.145318,(((Cynocephalus_volans:0.015408,Galeopterus_variegatus:0.020258)'1.000000':0.179481,(Tupaia_glis:0.064054,Ptilocercus_lowii:0.073474)'1.000000':0.061278)'0.725000':0.039329,((Lemur_catta:0.08319,Notharctus_tenebrosus:0.105118)'0.502778':0.029059,(Tarsius_syrichta:0.141359,(Homo_sapiens:0.157689,Saimiri_sciureus:0.070453)'0.994444':0.052494)'0.811111':0.027917)'1.000000':0.066765)'0.794444':0.048991)'0.644444':0.030381)'0.127778':0.020166)'0.502778':0.036002,((Solenodon_paradoxus:0.186659,((Erinaceus_europaeus:0.126378,Echinops_telfairi:0.145324)'0.941667':0.040873,(Amblysomus_hottentotus:0.226303,(Talpa_europaea:0.137076,Sorex_araneus:0.140941)'0.736111':0.05774)'0.961111':0.065367)'0.638889':0.027032)'0.800000':0.034082,(Leptictis_dakotensis:0.13621,Rhynchocyon_cirnei:0.16323)'0.872222':0.053568)'0.644444':0.037501)'0.944444':0.046542)'0.455556':0.035666)'0.736111':0.045448)'1.000000':0.073351,(Zhangheotherium_quinquecuspidens:0.057829,Henkelotherium_guimarotae:0.066552)'0.847222':0.040923)'1.000000':0.060415,(Ornithorhynchus_anatinus:0.080608,Tachyglossus_aculeatus:0.087246)'1.000000':0.137233)'1.000000':0.062624)'1.000000':0.077294):0);

[no PAs: nodal values = "transfer bootstrap expectation" values]
(Morganucodon_oehleri:0.007443,(Morganucodon_watsoni:0.008777,(Haldanodon_exspectatus:0.099164,(((Eomaia_scansoria:0.06223,((Dromiciops_gliroides:0.084822,(Pucadelphys_andinus:0.114211,Didelphis_virginiana:0.053163)'0.936111':0.054776)'1.000000':0.073516,((Maelestes_gobiensis:0.103069,(Zalambdalestes_lechei:0.124749,Ukhaatherium_nessovi:0.103981)'0.983333':0.055483)'0.811111':0.043684,((((Sinopa_rapax:0.089923,((Vulpavus_ovatus:0.017136,Vulpavus_profectus:0.02736)'1.000000':0.054969,(Felis_silvestris:0.09397,Canis_lupus:0.068977)'1.000000':0.061354)'0.962037':0.030009)'1.000000':0.077075,(Protungulatum_donnae:0.135793,((((Cocomys_lingchaensis:0.074623,Paramys_delicatus:0.05218)'0.355556':0.018293,(Rattus_norvegicus:0.151467,(Ictidomys_tridecemlineatus:0.063015,(Cavia_porcellus:0.090784,Castor_canadensis:0.086457)'0.947222':0.049951)'0.798611':0.0461)'0.762963':0.027639)'0.910556':0.033041,(Tribosphenomys_minutus:0.104868,(Rhombomylus_turpanensis:0.074478,(Gomphos_elkema:0.077086,(Dawsonolagus_antiquus:0.025474,Oryctolagus_cuniculus:0.098581)'0.975000':0.035809)'1.000000':0.072567)'0.872222':0.024996)'0.911111':0.033814)'1.000000':0.095354,((Apheliscus_insidiosus:0.114763,((Didolodus_multicuspis:0.070325,Protolipterna_ellipsodontoides:0.057358)'0.600000':0.033642,(Hyopsodus_paulus:0.100402,Phenacodus_intermedius:0.092737)'0.650000':0.026091)'0.737963':0.015702)'0.914583':0.026464,(((Mesonyx_obtusidens:0.155465,(Archaeotherium_mortoni:0.125507,(Rodhocetus_balochistanensis:0.104617,(Artiocetus_clavis:0.118052,(Basilosaurus_cetoides:0.141697,(Tursiops_truncatus:0.165577,Caperea_marginata:0.167985)'1.000000':0.113237)'0.791667':0.048819)'0.968519':0.051828)'0.886806':0.049302)'0.751667':0.026003)'0.782407':0.046651,((Equus_caballus:0.1459,Mesohippus_bairdi:0.151273)'0.866667':0.059552,((Hippopotamus_amphibius:0.19223,Sus_scrofa:0.160792)'0.225000':0.037404,(Lama_glama:0.121042,Bos_taurus:0.126859)'0.991667':0.060059)'0.512963':0.032286)'0.643333':0.044061)'0.957176':0.049429,((Procavia_capensis:0.194301,Thomashuxleya_externa_comb:0.13528)'0.302778':0.029765,(Carodnia_vieirai:0.087573,(Moeritherium_lyonsi:0.136601,(Trichechus_manatus:0.202685,Loxodonta_africana:0.168062)'0.505556':0.037237)'0.823611':0.042398)'0.983333':0.057215)'0.789444':0.04498)'0.961111':0.041203)'0.873068':0.035076)'0.904167':0.022444)'0.906111':0.019555)'0.892569':0.032656,((Orycteropus_afer:0.239321,((Metacheiromys_marshi:0.114432,Manis_pentadactyla:0.191407)'0.955556':0.059411,(Dasypus_novemcinctus:0.188778,(Hapalops_elongatus:0.157304,Tamandua_tetradactyla:0.142909)'1.000000':0.075132)'0.981944':0.055729)'0.984028':0.043583)'0.991667':0.06214,((Pteropus_giganteus:0.158564,((Onychonycteris_finneyi:0.031028,Icaronycteris_index:0.026973)'0.997222':0.060668,((Nycteris_thebaica:0.091164,(Rhinopoma_hardwickii:0.052008,Saccopteryx_bilineata:0.0521)'0.830556':0.025888)'0.993056':0.032396,(Myotis_lucifugus:0.061787,Pteronotus_parnellii:0.0761)'0.766667':0.026288)'1.000000':0.05286)'0.999074':0.05458)'1.000000':0.145318,(((Cynocephalus_volans:0.015408,Galeopterus_variegatus:0.020258)'1.000000':0.179481,(Tupaia_glis:0.064054,Ptilocercus_lowii:0.073474)'1.000000':0.061278)'0.816667':0.039329,((Lemur_catta:0.08319,Notharctus_tenebrosus:0.105118)'0.502778':0.029059,(Tarsius_syrichta:0.141359,(Homo_sapiens:0.157689,Saimiri_sciureus:0.070453)'0.994444':0.052494)'0.902778':0.027917)'1.000000':0.066765)'0.926389':0.048991)'0.824826':0.030381)'0.646717':0.020166)'0.855556':0.036002,((Solenodon_paradoxus:0.186659,((Erinaceus_europaeus:0.126378,Echinops_telfairi:0.145324)'0.941667':0.040873,(Amblysomus_hottentotus:0.226303,(Talpa_europaea:0.137076,Sorex_araneus:0.140941)'0.736111':0.05774)'0.980556':0.065367)'0.880556':0.027032)'0.927778':0.034082,(Leptictis_dakotensis:0.13621,Rhynchocyon_cirnei:0.16323)'0.872222':0.053568)'0.878968':0.037501)'0.988889':0.046542)'0.866389':0.035666)'0.962302':0.045448)'1.000000':0.073351,(Zhangheotherium_quinquecuspidens:0.057829,Henkelotherium_guimarotae:0.066552)'0.847222':0.040923)'1.000000':0.060415,(Ornithorhynchus_anatinus:0.080608,Tachyglossus_aculeatus:0.087246)'1.000000':0.137233)'1.000000':0.062624)'1.000000':0.077294):0);

["all characters" PAs corrected for the O'Leary et al. character ontology: nodal values = standard bootstrap values]
(Morganucodon_watsoni:0.007288,((((Tachyglossus_aculeatus:0.075508,(Ornithorhynchus_anatinus:0.000001,PA2:0.000001)1.000000:0.075106)1.000000:0.065036,(((((((Solenodon_paradoxus:0.106273,(PA23:0.000001,((Erinaceus_europaeus:0.094966,PA24:0.000001)0.911111:0.054204,(Talpa_europaea:0.108471,(PA25:0.000001,Sorex_araneus:0.000001)1.000000:0.056519)1.000000:0.077604)0.947222:0.057487)0.922222:0.046756)1.000000:0.085157,(((Manis_pentadactyla:0.168002,Metacheiromys_marshi:0.144333)0.991667:0.106640,((((Vulpavus_profectus:0.026430,Vulpavus_ovatus:0.016511)1.000000:0.050962,Sinopa_rapax:0.110294)0.991667:0.076829,(Canis_lupus:0.059151,((Felis_silvestris:0.000001,PA22:0.000001)1.000000:0.026347,PA21:0.000001)1.000000:0.046527)0.969444:0.037006)0.938889:0.055454,(((PA9:0.004871,(((Lama_glama:0.083254,(PA11:0.000001,(Sus_scrofa:0.118362,(((PA14:0.000001,Hippopotamus_amphibius:0.000001)1.000000:0.172795,((((Tursiops_truncatus:0.000001,PA15:0.000001)1.000000:0.120438,Caperea_marginata:0.154688)1.000000:0.230730,PA13:0.000001)0.183333:0.010596,Bos_taurus:0.035348)0.425000:0.072324)0.338889:0.030275,PA12:0.000001)0.338889:0.035892)0.419444:0.053741)0.344444:0.038824)0.416667:0.047514,PA10:0.005296)0.419444:0.056152,(Mesohippus_bairdi:0.220516,Equus_caballus:0.077767)0.683333:0.044460)0.205556:0.041041)0.227778:0.091837,(Pteropus_giganteus:0.154036,(((PA20:0.000001,Rhinopoma_hardwickii:0.000001)1.000000:0.047356,((((PA17:0.000001,Myotis_lucifugus:0.053276)0.969444:0.009386,Pteronotus_parnellii:0.067446)0.750000:0.018371,((Nycteris_thebaica:0.089276,PA18:0.008435)0.788889:0.011406,(Saccopteryx_bilineata:0.000001,PA19:0.000001)1.000000:0.044915)1.000000:0.026147)0.875000:0.036422,PA16:0.000001)0.852778:0.021975)0.852778:0.036024,(Onychonycteris_finneyi:0.033258,Icaronycteris_index:0.022125)1.000000:0.120416)0.494444:0.020509)1.000000:0.121718)0.172222:0.063680,PA8:0.001837)0.152778:0.045653)0.172222:0.032473)0.102778:0.051096,PA7:0.000001)0.080556:0.040122)0.116667:0.052616,(((Ptilocercus_lowii:0.063687,(PA27:0.007556,(Tupaia_glis:0.000001,PA32:0.000001)1.000000:0.049636)0.991667:0.020511)1.000000:0.055899,(((PA33:0.000001,((PA35:0.007133,(Tarsius_syrichta:0.110739,(((Saimiri_sciureus:0.000001,PA37:0.000001)1.000000:0.044275,PA36:0.003650)0.913889:0.015890,Homo_sapiens:0.148801)1.000000:0.053598)0.808333:0.018482)0.905556:0.037127,(Lemur_catta:0.040177,Notharctus_tenebrosus:0.152922)0.861111:0.021229)0.933333:0.031496)1.000000:0.061643,((Cynocephalus_volans:0.000001,PA34:0.000001)1.000000:0.014770,Galeopterus_variegatus:0.017415)1.000000:0.139668)1.000000:0.065483,PA26:0.000001)0.952778:0.034439)1.000000:0.099708,PA6:0.000001)0.402778:0.027495)0.066667:0.032303,(PA5:0.004070,(((((Rhombomylus_turpanensis:0.081992,((Dawsonolagus_antiquus:0.025261,Oryctolagus_cuniculus:0.093014)0.975000:0.032116,Gomphos_elkema:0.079233)1.000000:0.061711)0.725000:0.020232,Tribosphenomys_minutus:0.113031)0.711111:0.028568,((Paramys_delicatus:0.051490,Cocomys_lingchaensis:0.072822)0.863889:0.028653,(Rattus_norvegicus:0.119869,(PA28:0.009922,(((PA29:0.000001,(PA30:0.000001,Castor_canadensis:0.000001)1.000000:0.083755)0.813889:0.021464,(Ictidomys_tridecemlineatus:0.000001,PA31:0.000001)1.000000:0.073214)0.447222:0.018013,Cavia_porcellus:0.079390)0.719444:0.012662)0.936111:0.058427)0.969444:0.051608)0.469444:0.022849)1.000000:0.102339,(((Leptictis_dakotensis:0.144577,(Zalambdalestes_lechei:0.126095,(Ukhaatherium_nessovi:0.105289,(Eomaia_scansoria:0.083657,Maelestes_gobiensis:0.076657)0.488889:0.052095)0.175000:0.031784)0.666667:0.065520)0.272222:0.032425,Protungulatum_donnae:0.129190)0.083333:0.036495,((Mesonyx_obtusidens:0.129780,(Archaeotherium_mortoni:0.116236,(Basilosaurus_cetoides:0.201028,(Artiocetus_clavis:0.135692,Rodhocetus_balochistanensis:0.091696)0.113889:0.006286)0.263889:0.065061)0.169444:0.041744)0.419444:0.088642,((Carodnia_vieirai:0.117563,Thomashuxleya_externa_comb:0.122540)0.638889:0.066702,(((Phenacodus_intermedius:0.091199,Hyopsodus_paulus:0.098551)0.455556:0.025418,(Didolodus_multicuspis:0.074104,Protolipterna_ellipsodontoides:0.053149)0.569444:0.029178)0.130556:0.012272,Apheliscus_insidiosus:0.116539)0.200000:0.017336)0.163889:0.023646)0.061111:0.039835)0.025000:0.033744)0.008333:0.040263,(((((Tamandua_tetradactyla:0.109972,Hapalops_elongatus:0.181406)1.000000:0.078542,(PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.150520)0.886111:0.088111,PA38:0.000001)0.680556:0.028980,((((PA42:0.000001,Procavia_capensis:0.000001)1.000000:0.088359,(Trichechus_manatus:0.208371,PA41:0.000001)0.416667:0.014513)0.519444:0.064569,(Moeritherium_lyonsi:0.180980,Loxodonta_africana:0.134920)0.447222:0.064609)0.527778:0.112933,PA40:0.008434)0.422222:0.030982)0.252778:0.028628,((PA43:0.000001,(Rhynchocyon_cirnei:0.078088,(PA44:0.000001,(Amblysomus_hottentotus:0.210027,(PA45:0.000001,Echinops_telfairi:0.000001)1.000000:0.134173)0.452778:0.027792)0.766667:0.031688)0.647222:0.040955)0.991667:0.050178,Orycteropus_afer:0.165822)0.941667:0.043044)0.388889:0.088077)0.013889:0.037714)0.005556:0.021274)0.272222:0.045119,PA3:0.000001)0.275000:0.036609,((Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.053501,(Pucadelphys_andinus:0.129208,Didelphis_virginiana:0.035498)1.000000:0.055197)1.000000:0.093106)0.475000:0.074294,PA1:0.000001)0.475000:0.061732)0.997222:0.098712,(Zhangheotherium_quinquecuspidens:0.054445,Henkelotherium_guimarotae:0.078583)0.936111:0.060097)1.000000:0.103245,Haldanodon_exspectatus:0.096681)1.000000:0.077080,Morganucodon_oehleri:0.009027);

["all characters" PAs corrected for the O'Leary et al. character ontology: nodal values = "transfer bootstrap expectation" values]
(Morganucodon_watsoni:0.007288,((((Tachyglossus_aculeatus:0.075508,(Ornithorhynchus_anatinus:0.000001,PA2:0.000001)1.000000:0.075106)1.000000:0.065036,(((((((Solenodon_paradoxus:0.106273,(PA23:0.000001,((Erinaceus_europaeus:0.094966,PA24:0.000001)0.911111:0.054204,(Talpa_europaea:0.108471,(PA25:0.000001,Sorex_araneus:0.000001)1.000000:0.056519)1.000000:0.077604)0.974306:0.057487)0.984444:0.046756)1.000000:0.085157,(((Manis_pentadactyla:0.168002,Metacheiromys_marshi:0.144333)0.991667:0.106640,((((Vulpavus_profectus:0.026430,Vulpavus_ovatus:0.016511)1.000000:0.050962,Sinopa_rapax:0.110294)0.995833:0.076829,(Canis_lupus:0.059151,((Felis_silvestris:0.000001,PA22:0.000001)1.000000:0.026347,PA21:0.000001)1.000000:0.046527)0.989815:0.037006)0.969444:0.055454,(((PA9:0.004871,(((Lama_glama:0.083254,(PA11:0.000001,(Sus_scrofa:0.118362,(((PA14:0.000001,Hippopotamus_amphibius:0.000001)1.000000:0.172795,((((Tursiops_truncatus:0.000001,PA15:0.000001)1.000000:0.120438,Caperea_marginata:0.154688)1.000000:0.230730,PA13:0.000001)0.727778:0.010596,Bos_taurus:0.035348)0.715278:0.072324)0.653704:0.030275,PA12:0.000001)0.673810:0.035892)0.730208:0.053741)0.736728:0.038824)0.783889:0.047514,PA10:0.005296)0.804040:0.056152,(Mesohippus_bairdi:0.220516,Equus_caballus:0.077767)0.683333:0.044460)0.808120:0.041041)0.825992:0.091837,(Pteropus_giganteus:0.154036,(((PA20:0.000001,Rhinopoma_hardwickii:0.000001)1.000000:0.047356,((((PA17:0.000001,Myotis_lucifugus:0.053276)0.969444:0.009386,Pteronotus_parnellii:0.067446)0.859722:0.018371,((Nycteris_thebaica:0.089276,PA18:0.008435)0.788889:0.011406,(Saccopteryx_bilineata:0.000001,PA19:0.000001)1.000000:0.044915)1.000000:0.026147)0.958333:0.036422,PA16:0.000001)0.961111:0.021975)0.967901:0.036024,(Onychonycteris_finneyi:0.033258,Icaronycteris_index:0.022125)1.000000:0.120416)0.954040:0.020509)1.000000:0.121718)0.813683:0.063680,PA8:0.001837)0.823413:0.045653)0.887222:0.032473)0.875676:0.051096,PA7:0.000001)0.867032:0.040122)0.902840:0.052616,(((Ptilocercus_lowii:0.063687,(PA27:0.007556,(Tupaia_glis:0.000001,PA32:0.000001)1.000000:0.049636)0.995833:0.020511)1.000000:0.055899,(((PA33:0.000001,((PA35:0.007133,(Tarsius_syrichta:0.110739,(((Saimiri_sciureus:0.000001,PA37:0.000001)1.000000:0.044275,PA36:0.003650)0.956944:0.015890,Homo_sapiens:0.148801)1.000000:0.053598)0.952083:0.018482)0.963889:0.037127,(Lemur_catta:0.040177,Notharctus_tenebrosus:0.152922)0.861111:0.021229)0.990476:0.031496)1.000000:0.061643,((Cynocephalus_volans:0.000001,PA34:0.000001)1.000000:0.014770,Galeopterus_variegatus:0.017415)1.000000:0.139668)1.000000:0.065483,PA26:0.000001)0.996065:0.034439)1.000000:0.099708,PA6:0.000001)0.964869:0.027495)0.799559:0.032303,(PA5:0.004070,(((((Rhombomylus_turpanensis:0.081992,((Dawsonolagus_antiquus:0.025261,Oryctolagus_cuniculus:0.093014)0.975000:0.032116,Gomphos_elkema:0.079233)1.000000:0.061711)0.908333:0.020232,Tribosphenomys_minutus:0.113031)0.923611:0.028568,((Paramys_delicatus:0.051490,Cocomys_lingchaensis:0.072822)0.863889:0.028653,(Rattus_norvegicus:0.119869,(PA28:0.009922,(((PA29:0.000001,(PA30:0.000001,Castor_canadensis:0.000001)1.000000:0.083755)0.906944:0.021464,(Ictidomys_tridecemlineatus:0.000001,PA31:0.000001)1.000000:0.073214)0.809722:0.018013,Cavia_porcellus:0.079390)0.932778:0.012662)0.987963:0.058427)0.994048:0.051608)0.883333:0.022849)1.000000:0.102339,(((Leptictis_dakotensis:0.144577,(Zalambdalestes_lechei:0.126095,(Ukhaatherium_nessovi:0.105289,(Eomaia_scansoria:0.083657,Maelestes_gobiensis:0.076657)0.488889:0.052095)0.427778:0.031784)0.876852:0.065520)0.750694:0.032425,Protungulatum_donnae:0.129190)0.623889:0.036495,((Mesonyx_obtusidens:0.129780,(Archaeotherium_mortoni:0.116236,(Basilosaurus_cetoides:0.201028,(Artiocetus_clavis:0.135692,Rodhocetus_balochistanensis:0.091696)0.113889:0.006286)0.366667:0.065061)0.374074:0.041744)0.610417:0.088642,((Carodnia_vieirai:0.117563,Thomashuxleya_externa_comb:0.122540)0.638889:0.066702,(((Phenacodus_intermedius:0.091199,Hyopsodus_paulus:0.098551)0.455556:0.025418,(Didolodus_multicuspis:0.074104,Protolipterna_ellipsodontoides:0.053149)0.569444:0.029178)0.516667:0.012272,Apheliscus_insidiosus:0.116539)0.675000:0.017336)0.743981:0.023646)0.616667:0.039835)0.599020:0.033744)0.646788:0.040263,(((((Tamandua_tetradactyla:0.109972,Hapalops_elongatus:0.181406)1.000000:0.078542,(PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.150520)0.925000:0.088111,PA38:0.000001)0.889583:0.028980,((((PA42:0.000001,Procavia_capensis:0.000001)1.000000:0.088359,(Trichechus_manatus:0.208371,PA41:0.000001)0.416667:0.014513)0.836111:0.064569,(Moeritherium_lyonsi:0.180980,Loxodonta_africana:0.134920)0.447222:0.064609)0.903889:0.112933,PA40:0.008434)0.890278:0.030982)0.736364:0.028628,((PA43:0.000001,(Rhynchocyon_cirnei:0.078088,(PA44:0.000001,(Amblysomus_hottentotus:0.210027,(PA45:0.000001,Echinops_telfairi:0.000001)1.000000:0.134173)0.726389:0.027792)0.910185:0.031688)0.911111:0.040955)0.998333:0.050178,Orycteropus_afer:0.165822)0.990278:0.043044)0.940741:0.088077)0.688725:0.037714)0.688141:0.021274)0.787821:0.045119,PA3:0.000001)0.771065:0.036609,((Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.053501,(Pucadelphys_andinus:0.129208,Didelphis_virginiana:0.035498)1.000000:0.055197)1.000000:0.093106)0.754167:0.074294,PA1:0.000001)0.783730:0.061732)0.998611:0.098712,(Zhangheotherium_quinquecuspidens:0.054445,Henkelotherium_guimarotae:0.078583)0.936111:0.060097)1.000000:0.103245,Haldanodon_exspectatus:0.096681)1.000000:0.077080,Morganucodon_oehleri:0.009027);

["all characters" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included nodal values = standard bootstrap values]
(Morganucodon_watsoni:0.007321,(((Henkelotherium_guimarotae:0.078864,Zhangheotherium_quinquecuspidens:0.054664)0.944444:0.060414,((((PA3:0.000001,(((((Zalambdalestes_lechei:0.121713,((Eomaia_scansoria:0.081117,Maelestes_gobiensis:0.081313)0.491667:0.049625,Ukhaatherium_nessovi:0.108201)0.169444:0.037899)0.686111:0.066303,Leptictis_dakotensis:0.143033)0.238889:0.039656,((Apheliscus_insidiosus:0.108917,((Protolipterna_ellipsodontoides:0.052944,Didolodus_multicuspis:0.069148)0.602778:0.028497,((Phenacodus_intermedius:0.089382,Hyopsodus_paulus:0.099801)0.458333:0.026577,(Thomashuxleya_externa_comb:0.122565,Carodnia_vieirai:0.122471)0.611111:0.071595)0.086111:0.014127)0.125000:0.019870)0.258333:0.028654,Protungulatum_donnae:0.119696)0.355556:0.037908)0.083333:0.054663,(((PA43:0.000001,(Echinops_telfairi:0.104802,(Rhynchocyon_cirnei:0.113238,(Amblysomus_hottentotus:0.238042,PA44:0.000001)0.650000:0.000001)0.455556:0.036527)0.341667:0.036551)0.983333:0.046443,Orycteropus_afer:0.169796)0.936111:0.042047,((((PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.150489,(Tamandua_tetradactyla:0.109870,Hapalops_elongatus:0.181895)0.997222:0.078554)0.880556:0.088096,PA38:0.000001)0.694444:0.028539,((((PA41:0.000001,(Procavia_capensis:0.000001,PA42:0.000001)1.000000:0.104237)0.550000:0.040522,Trichechus_manatus:0.174211)0.494444:0.039656,(Loxodonta_africana:0.131844,Moeritherium_lyonsi:0.188079)0.397222:0.064558)0.511111:0.110461,PA40:0.008666)0.411111:0.031209)0.222222:0.032502)0.375000:0.081015)0.052778:0.033144,(PA5:0.009465,((((Rattus_norvegicus:0.140632,(Ictidomys_tridecemlineatus:0.078888,((Castor_canadensis:0.092894,Cavia_porcellus:0.083791)0.666667:0.012652,PA28:0.000001)0.825000:0.029248)0.808333:0.035868)0.869444:0.037115,(((Rhombomylus_turpanensis:0.085811,((Oryctolagus_cuniculus:0.090222,Dawsonolagus_antiquus:0.023507)0.980556:0.030309,Gomphos_elkema:0.081664)1.000000:0.059535)0.700000:0.022801,Tribosphenomys_minutus:0.104601)0.725000:0.028757,(Cocomys_lingchaensis:0.072292,Paramys_delicatus:0.052820)0.786111:0.034986)0.202778:0.025127)1.000000:0.120560,((Ptilocercus_lowii:0.064199,(Tupaia_glis:0.051752,PA27:0.004244)0.991667:0.021316)1.000000:0.064927,((((Saimiri_sciureus:0.066329,Homo_sapiens:0.156962)1.000000:0.055209,(((Tarsius_syrichta:0.128468,Notharctus_tenebrosus:0.118048)0.647222:0.044887,Lemur_catta:0.066598)0.558333:0.023646,PA33:0.000001)0.555556:0.006179)1.000000:0.057387,(Galeopterus_variegatus:0.018822,Cynocephalus_volans:0.016409)1.000000:0.139696)1.000000:0.062069,PA26:0.000001)0.955556:0.025498)1.000000:0.054272)0.430556:0.040399,(((PA7:0.000001,(Solenodon_paradoxus:0.172241,(Erinaceus_europaeus:0.158188,(Talpa_europaea:0.149266,Sorex_araneus:0.129835)1.000000:0.071613)0.838889:0.036569)0.902778:0.046184)0.541667:0.038803,((Manis_pentadactyla:0.171627,Metacheiromys_marshi:0.143501)0.997222:0.112393,(((Canis_lupus:0.059573,(Felis_silvestris:0.030358,PA21:0.000001)1.000000:0.046183)0.961111:0.039483,(Sinopa_rapax:0.110151,(Vulpavus_ovatus:0.016426,Vulpavus_profectus:0.026611)1.000000:0.050570)0.997222:0.074303)0.963889:0.055553,(((((((Saccopteryx_bilineata:0.051493,Rhinopoma_hardwickii:0.050675)0.708333:0.020723,Nycteris_thebaica:0.096503)0.994444:0.028392,(Pteronotus_parnellii:0.074385,Myotis_lucifugus:0.061301)0.794444:0.026971)0.958333:0.029468,(Onychonycteris_finneyi:0.031308,Icaronycteris_index:0.025504)1.000000:0.095806)0.944444:0.038429,Pteropus_giganteus:0.178982)1.000000:0.101564,(PA9:0.005773,((Mesohippus_bairdi:0.217973,Equus_caballus:0.079542)0.838889:0.044375,(((Hippopotamus_amphibius:0.153147,(Archaeotherium_mortoni:0.118170,(Mesonyx_obtusidens:0.149255,(Rodhocetus_balochistanensis:0.100626,(Basilosaurus_cetoides:0.153381,((Caperea_marginata:0.162309,Tursiops_truncatus:0.157518)1.000000:0.100964,Artiocetus_clavis:0.107528)0.416667:0.039324)0.955556:0.057635)0.736111:0.036830)0.722222:0.045214)0.858333:0.056903)0.772222:0.061133,Sus_scrofa:0.151554)0.744444:0.042201,(PA10:0.000001,(Bos_taurus:0.139530,Lama_glama:0.106896)0.936111:0.023223)0.941667:0.019245)0.738889:0.043632)0.730556:0.039055)0.897222:0.090810)0.602778:0.062364,PA8:0.002915)0.519444:0.044771)0.652778:0.021522)0.400000:0.029142)0.461111:0.069824,PA6:0.010224)0.158333:0.026961)0.230556:0.032076)0.105556:0.024401)0.247222:0.036124)0.247222:0.036577,((PA4:0.000001,Dromiciops_gliroides:0.000001)1.000000:0.053500,(Pucadelphys_andinus:0.129236,Didelphis_virginiana:0.035471)1.000000:0.055067)1.000000:0.093038)0.438889:0.074168,PA1:0.000001)0.438889:0.058948,(Ornithorhynchus_anatinus:0.080395,Tachyglossus_aculeatus:0.083968)1.000000:0.059619)1.000000:0.099997)1.000000:0.103870,Haldanodon_exspectatus:0.097074)1.000000:0.077767,Morganucodon_oehleri:0.009086);

["all characters" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: nodal values = "transfer bootstrap expectation" values]
(Morganucodon_watsoni:0.007321,(((Henkelotherium_guimarotae:0.078864,Zhangheotherium_quinquecuspidens:0.054664)0.944444:0.060414,((((PA3:0.000001,(((((Zalambdalestes_lechei:0.121713,((Eomaia_scansoria:0.081117,Maelestes_gobiensis:0.081313)0.491667:0.049625,Ukhaatherium_nessovi:0.108201)0.436111:0.037899)0.885185:0.066303,Leptictis_dakotensis:0.143033)0.742361:0.039656,((Apheliscus_insidiosus:0.108917,((Protolipterna_ellipsodontoides:0.052944,Didolodus_multicuspis:0.069148)0.602778:0.028497,((Phenacodus_intermedius:0.089382,Hyopsodus_paulus:0.099801)0.458333:0.026577,(Thomashuxleya_externa_comb:0.122565,Carodnia_vieirai:0.122471)0.611111:0.071595)0.361111:0.014127)0.640000:0.019870)0.819907:0.028654,Protungulatum_donnae:0.119696)0.878968:0.037908)0.664120:0.054663,(((PA43:0.000001,(Echinops_telfairi:0.104802,(Rhynchocyon_cirnei:0.113238,(Amblysomus_hottentotus:0.238042,PA44:0.000001)0.650000:0.000001)0.718056:0.036527)0.780556:0.036551)0.995833:0.046443,Orycteropus_afer:0.169796)0.987222:0.042047,((((PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.150489,(Tamandua_tetradactyla:0.109870,Hapalops_elongatus:0.181895)0.997222:0.078554)0.920370:0.088096,PA38:0.000001)0.891667:0.028539,((((PA41:0.000001,(Procavia_capensis:0.000001,PA42:0.000001)1.000000:0.104237)0.775000:0.040522,Trichechus_manatus:0.174211)0.825926:0.039656,(Loxodonta_africana:0.131844,Moeritherium_lyonsi:0.188079)0.397222:0.064558)0.900000:0.110461,PA40:0.008666)0.886111:0.031209)0.723990:0.032502)0.936275:0.081015)0.677130:0.033144,(PA5:0.009465,((((Rattus_norvegicus:0.140632,(Ictidomys_tridecemlineatus:0.078888,((Castor_canadensis:0.092894,Cavia_porcellus:0.083791)0.666667:0.012652,PA28:0.000001)0.908333:0.029248)0.934259:0.035868)0.960417:0.037115,(((Rhombomylus_turpanensis:0.085811,((Oryctolagus_cuniculus:0.090222,Dawsonolagus_antiquus:0.023507)0.980556:0.030309,Gomphos_elkema:0.081664)1.000000:0.059535)0.900000:0.022801,Tribosphenomys_minutus:0.104601)0.929861:0.028757,(Cocomys_lingchaensis:0.072292,Paramys_delicatus:0.052820)0.786111:0.034986)0.697222:0.025127)1.000000:0.120560,((Ptilocercus_lowii:0.064199,(Tupaia_glis:0.051752,PA27:0.004244)0.991667:0.021316)1.000000:0.064927,((((Saimiri_sciureus:0.066329,Homo_sapiens:0.156962)1.000000:0.055209,(((Tarsius_syrichta:0.128468,Notharctus_tenebrosus:0.118048)0.647222:0.044887,Lemur_catta:0.066598)0.779167:0.023646,PA33:0.000001)0.738889:0.006179)1.000000:0.057387,(Galeopterus_variegatus:0.018822,Cynocephalus_volans:0.016409)1.000000:0.139696)1.000000:0.062069,PA26:0.000001)0.994444:0.025498)1.000000:0.054272)0.715338:0.040399,(((PA7:0.000001,(Solenodon_paradoxus:0.172241,(Erinaceus_europaeus:0.158188,(Talpa_europaea:0.149266,Sorex_araneus:0.129835)1.000000:0.071613)0.919444:0.036569)0.962037:0.046184)0.861806:0.038803,((Manis_pentadactyla:0.171627,Metacheiromys_marshi:0.143501)0.997222:0.112393,(((Canis_lupus:0.059573,(Felis_silvestris:0.030358,PA21:0.000001)1.000000:0.046183)0.980556:0.039483,(Sinopa_rapax:0.110151,(Vulpavus_ovatus:0.016426,Vulpavus_profectus:0.026611)1.000000:0.050570)0.998611:0.074303)0.978333:0.055553,(((((((Saccopteryx_bilineata:0.051493,Rhinopoma_hardwickii:0.050675)0.708333:0.020723,Nycteris_thebaica:0.096503)0.997222:0.028392,(Pteronotus_parnellii:0.074385,Myotis_lucifugus:0.061301)0.794444:0.026971)0.979861:0.029468,(Onychonycteris_finneyi:0.031308,Icaronycteris_index:0.025504)1.000000:0.095806)0.990741:0.038429,Pteropus_giganteus:0.178982)1.000000:0.101564,(PA9:0.005773,((Mesohippus_bairdi:0.217973,Equus_caballus:0.079542)0.838889:0.044375,(((Hippopotamus_amphibius:0.153147,(Archaeotherium_mortoni:0.118170,(Mesonyx_obtusidens:0.149255,(Rodhocetus_balochistanensis:0.100626,(Basilosaurus_cetoides:0.153381,((Caperea_marginata:0.162309,Tursiops_truncatus:0.157518)1.000000:0.100964,Artiocetus_clavis:0.107528)0.708333:0.039324)0.985185:0.057635)0.933333:0.036830)0.930000:0.045214)0.959722:0.056903)0.945238:0.061133,Sus_scrofa:0.151554)0.928472:0.042201,(PA10:0.000001,(Bos_taurus:0.139530,Lama_glama:0.106896)0.936111:0.023223)0.966667:0.019245)0.928283:0.043632)0.950427:0.039055)0.965675:0.090810)0.872222:0.062364,PA8:0.002915)0.878502:0.044771)0.932375:0.021522)0.915591:0.029142)0.930324:0.069824,PA6:0.010224)0.914414:0.026961)0.802146:0.032076)0.784173:0.024401)0.756250:0.036124)0.734091:0.036577,((PA4:0.000001,Dromiciops_gliroides:0.000001)1.000000:0.053500,(Pucadelphys_andinus:0.129236,Didelphis_virginiana:0.035471)1.000000:0.055067)1.000000:0.093038)0.767460:0.074168,PA1:0.000001)0.814352:0.058948,(Ornithorhynchus_anatinus:0.080395,Tachyglossus_aculeatus:0.083968)1.000000:0.059619)1.000000:0.099997)1.000000:0.103870,Haldanodon_exspectatus:0.097074)1.000000:0.077767,Morganucodon_oehleri:0.009086);

["all characters" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: nodal values = standard bootstrap values]
(Morganucodon_watsoni:0.008493,(Haldanodon_exspectatus:0.091359,((Henkelotherium_guimarotae:0.066539,Zhangheotherium_quinquecuspidens:0.060354)0.918860:0.043458,((Eomaia_scansoria:0.050900,((Zalambdalestes_lechei:0.130795,Ukhaatherium_nessovi:0.098224)0.817982:0.061414,Maelestes_gobiensis:0.094062)0.464912:0.039211)0.710526:0.042431,(((PA1:0.000001,(Ornithorhynchus_anatinus:0.084393,Tachyglossus_aculeatus:0.085656)1.000000:0.112931)0.504386:0.062331,((((Amblysomus_hottentotus:0.240576,Echinops_telfairi:0.123732)0.320175:0.057868,(((((PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.151923,(Hapalops_elongatus:0.183843,Tamandua_tetradactyla:0.110794)1.000000:0.079196)0.815789:0.088779,PA38:0.000001)0.695175:0.027205,(Orycteropus_afer:0.206882,(((Loxodonta_africana:0.189355,(Moeritherium_lyonsi:0.153725,Trichechus_manatus:0.160095)0.304825:0.046473)0.642544:0.085838,Procavia_capensis:0.186724)0.263158:0.042642,PA40:0.000149)0.258772:0.019834)0.243421:0.021663)0.197368:0.023381,Rhynchocyon_cirnei:0.140075)0.225877:0.046544)0.067982:0.068291,(PA5:0.010957,(((Leptictis_dakotensis:0.161400,Protungulatum_donnae:0.129554)0.151316:0.061840,((((((Protolipterna_ellipsodontoides:0.052759,Didolodus_multicuspis:0.074378)0.574561:0.034596,(Phenacodus_intermedius:0.089072,Hyopsodus_paulus:0.102201)0.432018:0.023251)0.210526:0.011502,Apheliscus_insidiosus:0.119891)0.377193:0.024139,(Carodnia_vieirai:0.119288,Thomashuxleya_externa_comb:0.128642)0.370614:0.061525)0.087719:0.041072,(Mesonyx_obtusidens:0.166574,(((Mesohippus_bairdi:0.150986,Equus_caballus:0.143403)0.978070:0.059500,((Bos_taurus:0.126753,Lama_glama:0.117557)0.993421:0.056609,(Sus_scrofa:0.173172,(((((Tursiops_truncatus:0.166919,Caperea_marginata:0.158568)1.000000:0.107834,Basilosaurus_cetoides:0.134319)0.594298:0.046548,Artiocetus_clavis:0.132957)0.956140:0.106821,Archaeotherium_mortoni:0.109046)0.168860:0.036176,Hippopotamus_amphibius:0.173000)0.190789:0.043278)0.103070:0.030666)0.100877:0.030095)0.131579:0.027609,Rodhocetus_balochistanensis:0.141527)0.245614:0.020055)0.953947:0.043991)0.081140:0.039446,(((Rattus_norvegicus:0.149298,(Ictidomys_tridecemlineatus:0.066873,(Castor_canadensis:0.085299,Cavia_porcellus:0.088696)0.958333:0.046334)0.493421:0.042479)0.313596:0.025947,(Cocomys_lingchaensis:0.075474,Paramys_delicatus:0.051022)0.403509:0.024937)0.646930:0.027533,(Tribosphenomys_minutus:0.112780,(Rhombomylus_turpanensis:0.079796,((Oryctolagus_cuniculus:0.099021,Dawsonolagus_antiquus:0.025507)0.973684:0.032765,Gomphos_elkema:0.079316)1.000000:0.069368)0.625000:0.023185)0.848684:0.034316)1.000000:0.088346)0.052632:0.025385)0.039474:0.038370,(((((Ptilocercus_lowii:0.075222,Tupaia_glis:0.061207)1.000000:0.052618,(((Notharctus_tenebrosus:0.131393,Lemur_catta:0.063130)0.844298:0.040038,((Homo_sapiens:0.154244,Saimiri_sciureus:0.071927)1.000000:0.048643,Tarsius_syrichta:0.144640)0.712719:0.024280)1.000000:0.073398,PA26:0.001572)0.517544:0.017292)0.896930:0.032297,(Cynocephalus_volans:0.014744,Galeopterus_variegatus:0.020693)1.000000:0.169025)1.000000:0.080785,PA6:0.002627)0.820175:0.027682,(((Pteropus_giganteus:0.179049,((Onychonycteris_finneyi:0.031401,Icaronycteris_index:0.026037)1.000000:0.079532,((Pteronotus_parnellii:0.074022,Myotis_lucifugus:0.062209)0.714912:0.026097,((Saccopteryx_bilineata:0.053344,Rhinopoma_hardwickii:0.049868)0.695175:0.023241,Nycteris_thebaica:0.095172)1.000000:0.030797)0.995614:0.040020)0.973684:0.038694)1.000000:0.140294,(((Erinaceus_europaeus:0.159563,(Talpa_europaea:0.150642,Sorex_araneus:0.131383)0.660088:0.071509)0.212719:0.035916,Solenodon_paradoxus:0.171746)0.241228:0.042262,PA7:0.008144)0.129386:0.016993)0.186404:0.031964,((Metacheiromys_marshi:0.120446,Manis_pentadactyla:0.194842)0.995614:0.105448,((Sinopa_rapax:0.107204,(Vulpavus_ovatus:0.017602,Vulpavus_profectus:0.025942)1.000000:0.053933)0.467105:0.040382,(Canis_lupus:0.072895,Felis_silvestris:0.088048)0.989035:0.042511)1.000000:0.063327)0.260965:0.030512)0.072368:0.039077)0.061404:0.021046)0.010965:0.022713)0.010965:0.026246)0.635965:0.028034,PA3:0.004773)0.635965:0.043340)0.434211:0.043906,((Pucadelphys_andinus:0.126122,Didelphis_virginiana:0.037127)1.000000:0.050757,(PA4:0.000001,Dromiciops_gliroides:0.000001)1.000000:0.056731)1.000000:0.067977)0.458333:0.065897)0.995614:0.079659)1.000000:0.115475)1.000000:0.081219,Morganucodon_oehleri:0.007724);

["all characters" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: nodal values = "transfer bootstrap expectation" values]
(Morganucodon_watsoni:0.008493,(Haldanodon_exspectatus:0.091359,((Henkelotherium_guimarotae:0.066539,Zhangheotherium_quinquecuspidens:0.060354)0.918860:0.043458,((Eomaia_scansoria:0.050900,((Zalambdalestes_lechei:0.130795,Ukhaatherium_nessovi:0.098224)0.817982:0.061414,Maelestes_gobiensis:0.094062)0.732456:0.039211)0.894737:0.042431,(((PA1:0.000001,(Ornithorhynchus_anatinus:0.084393,Tachyglossus_aculeatus:0.085656)1.000000:0.112931)0.752193:0.062331,((((Amblysomus_hottentotus:0.240576,Echinops_telfairi:0.123732)0.320175:0.057868,(((((PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.151923,(Hapalops_elongatus:0.183843,Tamandua_tetradactyla:0.110794)1.000000:0.079196)0.877924:0.088779,PA38:0.000001)0.876096:0.027205,(Orycteropus_afer:0.206882,(((Loxodonta_africana:0.189355,(Moeritherium_lyonsi:0.153725,Trichechus_manatus:0.160095)0.304825:0.046473)0.817982:0.085838,Procavia_capensis:0.186724)0.701023:0.042642,PA40:0.000149)0.645285:0.019834)0.625439:0.021663)0.751096:0.023381,Rhynchocyon_cirnei:0.140075)0.808214:0.046544)0.731275:0.068291,(PA5:0.010957,(((Leptictis_dakotensis:0.161400,Protungulatum_donnae:0.129554)0.151316:0.061840,((((((Protolipterna_ellipsodontoides:0.052759,Didolodus_multicuspis:0.074378)0.574561:0.034596,(Phenacodus_intermedius:0.089072,Hyopsodus_paulus:0.102201)0.432018:0.023251)0.601608:0.011502,Apheliscus_insidiosus:0.119891)0.823465:0.024139,(Carodnia_vieirai:0.119288,Thomashuxleya_externa_comb:0.128642)0.370614:0.061525)0.713450:0.041072,(Mesonyx_obtusidens:0.166574,(((Mesohippus_bairdi:0.150986,Equus_caballus:0.143403)0.978070:0.059500,((Bos_taurus:0.126753,Lama_glama:0.117557)0.993421:0.056609,(Sus_scrofa:0.173172,(((((Tursiops_truncatus:0.166919,Caperea_marginata:0.158568)1.000000:0.107834,Basilosaurus_cetoides:0.134319)0.797149:0.046548,Artiocetus_clavis:0.132957)0.982456:0.106821,Archaeotherium_mortoni:0.109046)0.781250:0.036176,Hippopotamus_amphibius:0.173000)0.712719:0.043278)0.639985:0.030666)0.637884:0.030095)0.839254:0.027609,Rodhocetus_balochistanensis:0.141527)0.918660:0.020055)0.987025:0.043991)0.837373:0.039446,(((Rattus_norvegicus:0.149298,(Ictidomys_tridecemlineatus:0.066873,(Castor_canadensis:0.085299,Cavia_porcellus:0.088696)0.958333:0.046334)0.746711:0.042479)0.665936:0.025947,(Cocomys_lingchaensis:0.075474,Paramys_delicatus:0.051022)0.403509:0.024937)0.856140:0.027533,(Tribosphenomys_minutus:0.112780,(Rhombomylus_turpanensis:0.079796,((Oryctolagus_cuniculus:0.099021,Dawsonolagus_antiquus:0.025507)0.973684:0.032765,Gomphos_elkema:0.079316)1.000000:0.069368)0.875000:0.023185)0.956140:0.034316)1.000000:0.088346)0.803947:0.025385)0.816064:0.038370,(((((Ptilocercus_lowii:0.075222,Tupaia_glis:0.061207)1.000000:0.052618,(((Notharctus_tenebrosus:0.131393,Lemur_catta:0.063130)0.844298:0.040038,((Homo_sapiens:0.154244,Saimiri_sciureus:0.071927)1.000000:0.048643,Tarsius_syrichta:0.144640)0.856360:0.024280)1.000000:0.073398,PA26:0.001572)0.903509:0.017292)0.970551:0.032297,(Cynocephalus_volans:0.014744,Galeopterus_variegatus:0.020693)1.000000:0.169025)1.000000:0.080785,PA6:0.002627)0.982018:0.027682,(((Pteropus_giganteus:0.179049,((Onychonycteris_finneyi:0.031401,Icaronycteris_index:0.026037)1.000000:0.079532,((Pteronotus_parnellii:0.074022,Myotis_lucifugus:0.062209)0.714912:0.026097,((Saccopteryx_bilineata:0.053344,Rhinopoma_hardwickii:0.049868)0.695175:0.023241,Nycteris_thebaica:0.095172)1.000000:0.030797)0.997807:0.040020)0.995614:0.038694)1.000000:0.140294,(((Erinaceus_europaeus:0.159563,(Talpa_europaea:0.150642,Sorex_araneus:0.131383)0.660088:0.071509)0.456140:0.035916,Solenodon_paradoxus:0.171746)0.527778:0.042262,PA7:0.008144)0.467105:0.016993)0.783808:0.031964,((Metacheiromys_marshi:0.120446,Manis_pentadactyla:0.194842)0.995614:0.105448,((Sinopa_rapax:0.107204,(Vulpavus_ovatus:0.017602,Vulpavus_profectus:0.025942)1.000000:0.053933)0.733553:0.040382,(Canis_lupus:0.072895,Felis_silvestris:0.088048)0.989035:0.042511)1.000000:0.063327)0.753655:0.030512)0.789935:0.039077)0.777558:0.021046)0.597128:0.022713)0.605117:0.026246)0.913925:0.028034,PA3:0.004773)0.908626:0.043340)0.752558:0.043906,((Pucadelphys_andinus:0.126122,Didelphis_virginiana:0.037127)1.000000:0.050757,(PA4:0.000001,Dromiciops_gliroides:0.000001)1.000000:0.056731)1.000000:0.067977)0.739857:0.065897)0.998355:0.079659)1.000000:0.115475)1.000000:0.081219,Morganucodon_oehleri:0.007724);

["skeletal only" PAs corrected for the O'Leary et al. character ontology: nodal values = standard bootstrap values]
((Haldanodon_exspectatus:0.097223,((Henkelotherium_guimarotae:0.079154,Zhangheotherium_quinquecuspidens:0.054702)0.936667:0.060446,((PA1:0.000001,(((Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.061003,(Pucadelphys_andinus:0.128832,Didelphis_virginiana:0.039677)1.000000:0.060572)1.000000:0.085565,((((((PA23:0.000001,((Erinaceus_europaeus:0.103001,PA24:0.000001)0.616667:0.051542,(Talpa_europaea:0.123743,(Sorex_araneus:0.000001,PA25:0.000001)1.000000:0.066157)1.000000:0.063942)0.966667:0.058627)0.910000:0.045217,Solenodon_paradoxus:0.112646)1.000000:0.082479,(PA7:0.000723,((Metacheiromys_marshi:0.143841,Manis_pentadactyla:0.171825)0.993333:0.108678,(((Sinopa_rapax:0.110182,(Vulpavus_ovatus:0.016992,Vulpavus_profectus:0.026106)1.000000:0.051654)0.953333:0.071774,(Canis_lupus:0.058502,(PA21:0.000001,(PA22:0.000001,Felis_silvestris:0.000001)1.000000:0.028778)1.000000:0.047077)0.966667:0.044068)0.970000:0.062692,((((Onychonycteris_finneyi:0.034259,Icaronycteris_index:0.021902)1.000000:0.108943,(((Rhinopoma_hardwickii:0.000001,PA20:0.000001)1.000000:0.050782,((((Nycteris_thebaica:0.090771,PA18:0.006209)0.906667:0.011465,(Saccopteryx_bilineata:0.000001,PA19:0.000001)1.000000:0.046191)1.000000:0.028064,(Pteronotus_parnellii:0.071539,(PA17:0.000001,Myotis_lucifugus:0.053423)0.993333:0.008749)0.843333:0.019614)0.893333:0.029999,PA16:0.000001)0.860000:0.015264)0.913333:0.045404,Pteropus_giganteus:0.172791)0.280000:0.024207)1.000000:0.113899,(PA9:0.006504,((Mesohippus_bairdi:0.218902,Equus_caballus:0.084022)0.623333:0.047952,((Lama_glama:0.087241,(PA11:0.000001,(((PA12:0.000001,Sus_scrofa:0.154400)0.473333:0.010444,(PA14:0.000001,Hippopotamus_amphibius:0.000001)1.000000:0.198174)0.366667:0.034341,(((Caperea_marginata:0.156183,(Tursiops_truncatus:0.000001,PA15:0.000001)1.000000:0.125139)1.000000:0.240026,PA13:0.000001)0.306667:0.015649,Bos_taurus:0.038253)0.380000:0.053470)0.356667:0.050377)0.320000:0.034833)0.353333:0.039766,PA10:0.008574)0.370000:0.054506)0.193333:0.033462)0.223333:0.093108)0.153333:0.050722,PA8:0.005083)0.153333:0.048001)0.156667:0.026023)0.070000:0.044381)0.036667:0.037457)0.083333:0.045273,(((Ptilocercus_lowii:0.065881,(PA27:0.007039,(Tupaia_glis:0.000001,PA32:0.000001)1.000000:0.047450)0.996667:0.021840)1.000000:0.054884,((((Cynocephalus_volans:0.000001,PA34:0.000001)1.000000:0.015827,Galeopterus_variegatus:0.019098)1.000000:0.143156,(PA33:0.000001,((Notharctus_tenebrosus:0.152876,Lemur_catta:0.042798)0.850000:0.022566,(PA35:0.009230,((Homo_sapiens:0.151257,(PA36:0.001934,(Saimiri_sciureus:0.000001,PA37:0.000001)1.000000:0.043883)0.753333:0.016146)1.000000:0.053629,Tarsius_syrichta:0.113933)0.616667:0.017476)0.860000:0.037516)0.883333:0.029777)1.000000:0.059532)0.986667:0.061644,PA26:0.000001)0.860000:0.031176)1.000000:0.095875,PA6:0.000001)0.713333:0.028253)0.073333:0.030686,((((((Rhombomylus_turpanensis:0.083354,((Dawsonolagus_antiquus:0.025248,Oryctolagus_cuniculus:0.095533)0.973333:0.033852,Gomphos_elkema:0.079194)1.000000:0.063237)0.666667:0.021226,Tribosphenomys_minutus:0.112773)0.790000:0.029946,((Paramys_delicatus:0.052178,Cocomys_lingchaensis:0.072848)0.756667:0.028387,(Rattus_norvegicus:0.118279,(PA28:0.009291,(((PA31:0.000001,Ictidomys_tridecemlineatus:0.000001)1.000000:0.074931,(PA29:0.000001,(PA30:0.000001,Castor_canadensis:0.000001)1.000000:0.072167)0.986667:0.022847)0.523333:0.024083,Cavia_porcellus:0.091924)0.196667:0.007037)0.913333:0.057877)0.956667:0.051737)0.670000:0.022776)1.000000:0.097598,(((Mesonyx_obtusidens:0.130203,((Rodhocetus_balochistanensis:0.098434,(Artiocetus_clavis:0.130695,Basilosaurus_cetoides:0.189989)0.320000:0.034599)0.223333:0.048830,Archaeotherium_mortoni:0.116193)0.163333:0.042035)0.366667:0.089701,((((Protolipterna_ellipsodontoides:0.053302,Didolodus_multicuspis:0.074005)0.576667:0.029139,(Hyopsodus_paulus:0.098888,Phenacodus_intermedius:0.091601)0.433333:0.025485)0.166667:0.012389,Apheliscus_insidiosus:0.116825)0.236667:0.016805,(Thomashuxleya_externa_comb:0.123972,Carodnia_vieirai:0.117433)0.616667:0.067735)0.193333:0.023603)0.053333:0.040196,(Protungulatum_donnae:0.130366,((Zalambdalestes_lechei:0.126879,(Ukhaatherium_nessovi:0.106018,(Maelestes_gobiensis:0.076856,Eomaia_scansoria:0.083762)0.480000:0.052368)0.176667:0.031686)0.663333:0.067043,Leptictis_dakotensis:0.144202)0.256667:0.032329)0.116667:0.036643)0.023333:0.032927)0.013333:0.038378,(((Orycteropus_afer:0.220852,((Trichechus_manatus:0.175193,(((PA42:0.000001,Procavia_capensis:0.000001)1.000000:0.104956,PA41:0.000016)0.720000:0.062431,(Moeritherium_lyonsi:0.189564,Loxodonta_africana:0.138397)0.386667:0.070971)0.176667:0.021880)0.483333:0.110553,PA40:0.002020)0.460000:0.018923)0.176667:0.013801,(((Dasypus_novemcinctus:0.000001,PA39:0.000001)1.000000:0.161092,(Hapalops_elongatus:0.181417,Tamandua_tetradactyla:0.117750)1.000000:0.083825)0.873333:0.084625,PA38:0.000001)0.530000:0.024282)0.193333:0.028527,(Rhynchocyon_cirnei:0.095846,(PA43:0.011917,((PA45:0.000001,Echinops_telfairi:0.000001)1.000000:0.129989,(Amblysomus_hottentotus:0.237180,PA44:0.000108)0.696667:0.029535)0.666667:0.032293)0.466667:0.026401)0.983333:0.049082)0.286667:0.078145)0.013333:0.035002,PA5:0.005504)0.010000:0.019863)0.270000:0.047728,PA3:0.000001)0.270000:0.034068)0.446667:0.068073)0.446667:0.070021,((PA2:0.000001,Ornithorhynchus_anatinus:0.000001)1.000000:0.076239,Tachyglossus_aculeatus:0.079739)1.000000:0.075057)0.993333:0.096850)1.000000:0.103543)1.000000:0.077603,Morganucodon_watsoni:0.007355,Morganucodon_oehleri:0.009061);

["skeletal only" PAs corrected for the O'Leary et al. character ontology: nodal values = "transfer bootstrap expectation" values]
((Haldanodon_exspectatus:0.097223,((Henkelotherium_guimarotae:0.079154,Zhangheotherium_quinquecuspidens:0.054702)0.936667:0.060446,((PA1:0.000001,(((Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.061003,(Pucadelphys_andinus:0.128832,Didelphis_virginiana:0.039677)1.000000:0.060572)1.000000:0.085565,((((((PA23:0.000001,((Erinaceus_europaeus:0.103001,PA24:0.000001)0.616667:0.051542,(Talpa_europaea:0.123743,(Sorex_araneus:0.000001,PA25:0.000001)1.000000:0.066157)1.000000:0.063942)0.985000:0.058627)0.982000:0.045217,Solenodon_paradoxus:0.112646)1.000000:0.082479,(PA7:0.000723,((Metacheiromys_marshi:0.143841,Manis_pentadactyla:0.171825)0.993333:0.108678,(((Sinopa_rapax:0.110182,(Vulpavus_ovatus:0.016992,Vulpavus_profectus:0.026106)1.000000:0.051654)0.976667:0.071774,(Canis_lupus:0.058502,(PA21:0.000001,(PA22:0.000001,Felis_silvestris:0.000001)1.000000:0.028778)1.000000:0.047077)0.988889:0.044068)0.985000:0.062692,((((Onychonycteris_finneyi:0.034259,Icaronycteris_index:0.021902)1.000000:0.108943,(((Rhinopoma_hardwickii:0.000001,PA20:0.000001)1.000000:0.050782,((((Nycteris_thebaica:0.090771,PA18:0.006209)0.906667:0.011465,(Saccopteryx_bilineata:0.000001,PA19:0.000001)1.000000:0.046191)1.000000:0.028064,(Pteronotus_parnellii:0.071539,(PA17:0.000001,Myotis_lucifugus:0.053423)0.993333:0.008749)0.918333:0.019614)0.961111:0.029999,PA16:0.000001)0.964762:0.015264)0.980741:0.045404,Pteropus_giganteus:0.172791)0.919333:0.024207)1.000000:0.113899,(PA9:0.006504,((Mesohippus_bairdi:0.218902,Equus_caballus:0.084022)0.623333:0.047952,((Lama_glama:0.087241,(PA11:0.000001,(((PA12:0.000001,Sus_scrofa:0.154400)0.473333:0.010444,(PA14:0.000001,Hippopotamus_amphibius:0.000001)1.000000:0.198174)0.585556:0.034341,(((Caperea_marginata:0.156183,(Tursiops_truncatus:0.000001,PA15:0.000001)1.000000:0.125139)1.000000:0.240026,PA13:0.000001)0.768889:0.015649,Bos_taurus:0.038253)0.695000:0.053470)0.723333:0.050377)0.734444:0.034833)0.775333:0.039766,PA10:0.008574)0.804848:0.054506)0.801282:0.033462)0.831667:0.093108)0.761481:0.050722,PA8:0.005083)0.777619:0.048001)0.840571:0.026023)0.829910:0.044381)0.826491:0.037457)0.878296:0.045273,(((Ptilocercus_lowii:0.065881,(PA27:0.007039,(Tupaia_glis:0.000001,PA32:0.000001)1.000000:0.047450)0.998333:0.021840)1.000000:0.054884,((((Cynocephalus_volans:0.000001,PA34:0.000001)1.000000:0.015827,Galeopterus_variegatus:0.019098)1.000000:0.143156,(PA33:0.000001,((Notharctus_tenebrosus:0.152876,Lemur_catta:0.042798)0.850000:0.022566,(PA35:0.009230,((Homo_sapiens:0.151257,(PA36:0.001934,(Saimiri_sciureus:0.000001,PA37:0.000001)1.000000:0.043883)0.876667:0.016146)1.000000:0.053629,Tarsius_syrichta:0.113933)0.904167:0.017476)0.946000:0.037516)0.983333:0.029777)1.000000:0.059532)0.996364:0.061644,PA26:0.000001)0.985000:0.031176)1.000000:0.095875,PA6:0.000001)0.983137:0.028253)0.858519:0.030686,((((((Rhombomylus_turpanensis:0.083354,((Dawsonolagus_antiquus:0.025248,Oryctolagus_cuniculus:0.095533)0.973333:0.033852,Gomphos_elkema:0.079194)1.000000:0.063237)0.888889:0.021226,Tribosphenomys_minutus:0.112773)0.945833:0.029946,((Paramys_delicatus:0.052178,Cocomys_lingchaensis:0.072848)0.756667:0.028387,(Rattus_norvegicus:0.118279,(PA28:0.009291,(((PA31:0.000001,Ictidomys_tridecemlineatus:0.000001)1.000000:0.074931,(PA29:0.000001,(PA30:0.000001,Castor_canadensis:0.000001)1.000000:0.072167)0.993333:0.022847)0.765833:0.024083,Cavia_porcellus:0.091924)0.822667:0.007037)0.985556:0.057877)0.993810:0.051737)0.929630:0.022776)1.000000:0.097598,(((Mesonyx_obtusidens:0.130203,((Rodhocetus_balochistanensis:0.098434,(Artiocetus_clavis:0.130695,Basilosaurus_cetoides:0.189989)0.320000:0.034599)0.328333:0.048830,Archaeotherium_mortoni:0.116193)0.340000:0.042035)0.558333:0.089701,((((Protolipterna_ellipsodontoides:0.053302,Didolodus_multicuspis:0.074005)0.576667:0.029139,(Hyopsodus_paulus:0.098888,Phenacodus_intermedius:0.091601)0.433333:0.025485)0.544444:0.012389,Apheliscus_insidiosus:0.116825)0.704167:0.016805,(Thomashuxleya_externa_comb:0.123972,Carodnia_vieirai:0.117433)0.616667:0.067735)0.740000:0.023603)0.590909:0.040196,(Protungulatum_donnae:0.130366,((Zalambdalestes_lechei:0.126879,(Ukhaatherium_nessovi:0.106018,(Maelestes_gobiensis:0.076856,Eomaia_scansoria:0.083762)0.480000:0.052368)0.423333:0.031686)0.871111:0.067043,Leptictis_dakotensis:0.144202)0.736667:0.032329)0.618667:0.036643)0.565098:0.032927)0.713125:0.038378,(((Orycteropus_afer:0.220852,((Trichechus_manatus:0.175193,(((PA42:0.000001,Procavia_capensis:0.000001)1.000000:0.104956,PA41:0.000016)0.860000:0.062431,(Moeritherium_lyonsi:0.189564,Loxodonta_africana:0.138397)0.386667:0.070971)0.717500:0.021880)0.896000:0.110553,PA40:0.002020)0.905556:0.018923)0.837619:0.013801,(((Dasypus_novemcinctus:0.000001,PA39:0.000001)1.000000:0.161092,(Hapalops_elongatus:0.181417,Tamandua_tetradactyla:0.117750)1.000000:0.083825)0.915556:0.084625,PA38:0.000001)0.842500:0.024282)0.767778:0.028527,(Rhynchocyon_cirnei:0.095846,(PA43:0.011917,((PA45:0.000001,Echinops_telfairi:0.000001)1.000000:0.129989,(Amblysomus_hottentotus:0.237180,PA44:0.000108)0.696667:0.029535)0.816667:0.032293)0.865833:0.026401)0.996667:0.049082)0.921667:0.078145)0.762614:0.035002,PA5:0.005504)0.763141:0.019863)0.783590:0.047728,PA3:0.000001)0.765556:0.034068)0.735417:0.068073)0.769048:0.070021,((PA2:0.000001,Ornithorhynchus_anatinus:0.000001)1.000000:0.076239,Tachyglossus_aculeatus:0.079739)1.000000:0.075057)0.997500:0.096850)1.000000:0.103543)1.000000:0.077603,Morganucodon_watsoni:0.007355,Morganucodon_oehleri:0.009061);

["skeletal only" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: nodal values = standard bootstrap values]
(Morganucodon_watsoni:0.008467,(((Henkelotherium_guimarotae:0.066795,Zhangheotherium_quinquecuspidens:0.059791)0.933333:0.043525,((Eomaia_scansoria:0.050236,(Maelestes_gobiensis:0.094196,(Ukhaatherium_nessovi:0.098072,Zalambdalestes_lechei:0.130355)0.808333:0.061103)0.488889:0.039613)0.672222:0.042300,(((Didelphis_virginiana:0.041249,Pucadelphys_andinus:0.124482)1.000000:0.055084,(PA4:0.000001,Dromiciops_gliroides:0.000001)1.000000:0.063798)1.000000:0.063155,(((((((Manis_pentadactyla:0.175595,Metacheiromys_marshi:0.140695)0.994444:0.111357,(((Canis_lupus:0.058727,(Felis_silvestris:0.033683,PA21:0.000001)1.000000:0.046386)0.975000:0.047877,((Vulpavus_profectus:0.026095,Vulpavus_ovatus:0.017072)1.000000:0.050954,Sinopa_rapax:0.110218)0.961111:0.068768)0.983333:0.064258,(((Pteropus_giganteus:0.177170,((Onychonycteris_finneyi:0.031404,Icaronycteris_index:0.025621)1.000000:0.094578,((Pteronotus_parnellii:0.075183,Myotis_lucifugus:0.061463)0.808333:0.026552,(Nycteris_thebaica:0.094983,(Saccopteryx_bilineata:0.051395,Rhinopoma_hardwickii:0.050927)0.747222:0.022246)0.997222:0.029517)0.977778:0.031224)0.936111:0.040791)1.000000:0.106703,(((Mesohippus_bairdi:0.217737,Equus_caballus:0.084751)0.719444:0.047155,(((Hippopotamus_amphibius:0.151876,(((Rodhocetus_balochistanensis:0.097476,(((Tursiops_truncatus:0.158905,Caperea_marginata:0.163662)1.000000:0.103444,Basilosaurus_cetoides:0.142983)0.555556:0.042906,Artiocetus_clavis:0.123943)0.941667:0.047432)0.686111:0.041374,Mesonyx_obtusidens:0.150889)0.588889:0.044729,Archaeotherium_mortoni:0.119335)0.727778:0.056882)0.613889:0.060392,Sus_scrofa:0.152627)0.547222:0.041899,((Bos_taurus:0.138511,Lama_glama:0.108534)0.875000:0.023494,PA10:0.000001)0.941667:0.021580)0.541667:0.042976)0.516667:0.034107,PA9:0.008256)0.911111:0.092136)0.369444:0.048577,PA8:0.006820)0.408333:0.046500)0.363889:0.018221)0.194444:0.027269,((((Sorex_araneus:0.130519,Talpa_europaea:0.150345)1.000000:0.072162,Erinaceus_europaeus:0.156856)0.736111:0.037172,Solenodon_paradoxus:0.173161)0.819444:0.045255,PA7:0.000753)0.530556:0.037228)0.252778:0.049191,(((Ptilocercus_lowii:0.065888,(PA27:0.005710,Tupaia_glis:0.048387)0.997222:0.022684)1.000000:0.055447,(PA26:0.000001,((Galeopterus_variegatus:0.019006,Cynocephalus_volans:0.016475)1.000000:0.143426,((Homo_sapiens:0.157795,Saimiri_sciureus:0.066727)1.000000:0.055982,(((Notharctus_tenebrosus:0.117938,Tarsius_syrichta:0.129585)0.658333:0.044958,Lemur_catta:0.067470)0.575000:0.026010,PA33:0.000001)0.544444:0.004265)1.000000:0.058486)0.994444:0.059256)0.844444:0.030953)1.000000:0.095873,PA6:0.000001)0.716667:0.026078)0.188889:0.032263,(PA5:0.003329,(((((Rhombomylus_turpanensis:0.084997,((Dawsonolagus_antiquus:0.025223,Oryctolagus_cuniculus:0.095872)0.980556:0.033650,Gomphos_elkema:0.079457)1.000000:0.061859)0.650000:0.020892,Tribosphenomys_minutus:0.113604)0.783333:0.029795,((Paramys_delicatus:0.052467,Cocomys_lingchaensis:0.072257)0.791667:0.029434,(Rattus_norvegicus:0.133003,(Ictidomys_tridecemlineatus:0.073109,((Cavia_porcellus:0.087452,Castor_canadensis:0.091680)0.725000:0.013139,PA28:0.000001)0.872222:0.035524)0.825000:0.041462)0.900000:0.037535)0.680556:0.025131)1.000000:0.098407,(Leptictis_dakotensis:0.180051,((Thomashuxleya_externa_comb:0.124765,Carodnia_vieirai:0.114780)0.602778:0.071236,((Hyopsodus_paulus:0.103492,Phenacodus_intermedius:0.088384)0.497222:0.024830,((Protolipterna_ellipsodontoides:0.050090,Didolodus_multicuspis:0.074646)0.586111:0.029050,(Apheliscus_insidiosus:0.101395,Protungulatum_donnae:0.136590)0.194444:0.031947)0.080556:0.018618)0.480556:0.031020)0.352778:0.038396)0.211111:0.040321)0.155556:0.036395,(((((PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.161014,(Tamandua_tetradactyla:0.117625,Hapalops_elongatus:0.181866)0.997222:0.083708)0.883333:0.084266,PA38:0.000001)0.530556:0.024135,(Orycteropus_afer:0.221187,((Trichechus_manatus:0.175528,((Moeritherium_lyonsi:0.190467,Loxodonta_africana:0.138521)0.394444:0.070799,(PA41:0.000070,(Procavia_capensis:0.000001,PA42:0.000001)1.000000:0.104772)0.744444:0.062428)0.186111:0.021623)0.502778:0.110196,PA40:0.002194)0.461111:0.018931)0.200000:0.013808)0.200000:0.028192,(Rhynchocyon_cirnei:0.096154,(((Amblysomus_hottentotus:0.237959,PA44:0.000001)0.830556:0.031585,Echinops_telfairi:0.115967)0.466667:0.031463,PA43:0.011013)0.383333:0.027382)0.986111:0.049439)0.283333:0.074894)0.141667:0.038588)0.116667:0.020030)0.661111:0.044305,PA3:0.005114)0.661111:0.041908,(PA1:0.000001,(Tachyglossus_aculeatus:0.084129,Ornithorhynchus_anatinus:0.081614)1.000000:0.136642)0.552778:0.054701)0.483333:0.041203)0.494444:0.067035)0.997222:0.079008)1.000000:0.115070,Haldanodon_exspectatus:0.091085)1.000000:0.080986,Morganucodon_oehleri:0.007715);

["skeletal only" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: nodal values = "transfer bootstrap expectation" values]
(Morganucodon_watsoni:0.008467,(((Henkelotherium_guimarotae:0.066795,Zhangheotherium_quinquecuspidens:0.059791)0.933333:0.043525,((Eomaia_scansoria:0.050236,(Maelestes_gobiensis:0.094196,(Ukhaatherium_nessovi:0.098072,Zalambdalestes_lechei:0.130355)0.808333:0.061103)0.744444:0.039613)0.881481:0.042300,(((Didelphis_virginiana:0.041249,Pucadelphys_andinus:0.124482)1.000000:0.055084,(PA4:0.000001,Dromiciops_gliroides:0.000001)1.000000:0.063798)1.000000:0.063155,(((((((Manis_pentadactyla:0.175595,Metacheiromys_marshi:0.140695)0.994444:0.111357,(((Canis_lupus:0.058727,(Felis_silvestris:0.033683,PA21:0.000001)1.000000:0.046386)0.987500:0.047877,((Vulpavus_profectus:0.026095,Vulpavus_ovatus:0.017072)1.000000:0.050954,Sinopa_rapax:0.110218)0.980556:0.068768)0.990000:0.064258,(((Pteropus_giganteus:0.177170,((Onychonycteris_finneyi:0.031404,Icaronycteris_index:0.025621)1.000000:0.094578,((Pteronotus_parnellii:0.075183,Myotis_lucifugus:0.061463)0.808333:0.026552,(Nycteris_thebaica:0.094983,(Saccopteryx_bilineata:0.051395,Rhinopoma_hardwickii:0.050927)0.747222:0.022246)0.998611:0.029517)0.988889:0.031224)0.989352:0.040791)1.000000:0.106703,(((Mesohippus_bairdi:0.217737,Equus_caballus:0.084751)0.719444:0.047155,(((Hippopotamus_amphibius:0.151876,(((Rodhocetus_balochistanensis:0.097476,(((Tursiops_truncatus:0.158905,Caperea_marginata:0.163662)1.000000:0.103444,Basilosaurus_cetoides:0.142983)0.777778:0.042906,Artiocetus_clavis:0.123943)0.980556:0.047432)0.920139:0.041374,Mesonyx_obtusidens:0.150889)0.891667:0.044729,Archaeotherium_mortoni:0.119335)0.920833:0.056882)0.894048:0.060392,Sus_scrofa:0.152627)0.856250:0.041899,((Bos_taurus:0.138511,Lama_glama:0.108534)0.875000:0.023494,PA10:0.000001)0.968056:0.021580)0.878283:0.042976)0.940385:0.034107,PA9:0.008256)0.972817:0.092136)0.800631:0.048577,PA8:0.006820)0.804589:0.046500)0.803448:0.018221)0.791129:0.027269,((((Sorex_araneus:0.130519,Talpa_europaea:0.150345)1.000000:0.072162,Erinaceus_europaeus:0.156856)0.868056:0.037172,Solenodon_paradoxus:0.173161)0.932407:0.045255,PA7:0.000753)0.849306:0.037228)0.831867:0.049191,(((Ptilocercus_lowii:0.065888,(PA27:0.005710,Tupaia_glis:0.048387)0.997222:0.022684)1.000000:0.055447,(PA26:0.000001,((Galeopterus_variegatus:0.019006,Cynocephalus_volans:0.016475)1.000000:0.143426,((Homo_sapiens:0.157795,Saimiri_sciureus:0.066727)1.000000:0.055982,(((Notharctus_tenebrosus:0.117938,Tarsius_syrichta:0.129585)0.658333:0.044958,Lemur_catta:0.067470)0.787500:0.026010,PA33:0.000001)0.744444:0.004265)1.000000:0.058486)0.998413:0.059256)0.979167:0.030953)1.000000:0.095873,PA6:0.000001)0.976389:0.026078)0.816043:0.032263,(PA5:0.003329,(((((Rhombomylus_turpanensis:0.084997,((Dawsonolagus_antiquus:0.025223,Oryctolagus_cuniculus:0.095872)0.980556:0.033650,Gomphos_elkema:0.079457)1.000000:0.061859)0.883333:0.020892,Tribosphenomys_minutus:0.113604)0.943750:0.029795,((Paramys_delicatus:0.052467,Cocomys_lingchaensis:0.072257)0.791667:0.029434,(Rattus_norvegicus:0.133003,(Ictidomys_tridecemlineatus:0.073109,((Cavia_porcellus:0.087452,Castor_canadensis:0.091680)0.725000:0.013139,PA28:0.000001)0.934722:0.035524)0.940741:0.041462)0.968750:0.037535)0.897685:0.025131)1.000000:0.098407,(Leptictis_dakotensis:0.180051,((Thomashuxleya_externa_comb:0.124765,Carodnia_vieirai:0.114780)0.602778:0.071236,((Hyopsodus_paulus:0.103492,Phenacodus_intermedius:0.088384)0.497222:0.024830,((Protolipterna_ellipsodontoides:0.050090,Didolodus_multicuspis:0.074646)0.586111:0.029050,(Apheliscus_insidiosus:0.101395,Protungulatum_donnae:0.136590)0.194444:0.031947)0.469444:0.018618)0.831111:0.031020)0.870238:0.038396)0.825000:0.040321)0.742222:0.036395,(((((PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.161014,(Tamandua_tetradactyla:0.117625,Hapalops_elongatus:0.181866)0.997222:0.083708)0.922222:0.084266,PA38:0.000001)0.848611:0.024135,(Orycteropus_afer:0.221187,((Trichechus_manatus:0.175528,((Moeritherium_lyonsi:0.190467,Loxodonta_africana:0.138521)0.394444:0.070799,(PA41:0.000070,(Procavia_capensis:0.000001,PA42:0.000001)1.000000:0.104772)0.872222:0.062428)0.729167:0.021623)0.900556:0.110196,PA40:0.002194)0.904167:0.018931)0.833730:0.013808)0.770139:0.028192,(Rhynchocyon_cirnei:0.096154,(((Amblysomus_hottentotus:0.237959,PA44:0.000001)0.830556:0.031585,Echinops_telfairi:0.115967)0.730556:0.031463,PA43:0.011013)0.793519:0.027382)0.996528:0.049439)0.916667:0.074894)0.716155:0.038588)0.714601:0.020030)0.923438:0.044305,PA3:0.005114)0.918333:0.041908,(PA1:0.000001,(Tachyglossus_aculeatus:0.084129,Ornithorhynchus_anatinus:0.081614)1.000000:0.136642)0.776389:0.054701)0.775694:0.041203)0.760417:0.067035)0.998611:0.079008)1.000000:0.115070,Haldanodon_exspectatus:0.091085)1.000000:0.080986,Morganucodon_oehleri:0.007715);

["skeletal only" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: nodal values = standard bootstrap values]
(((((Eomaia_scansoria:0.050666,(Maelestes_gobiensis:0.094692,(Ukhaatherium_nessovi:0.098999,Zalambdalestes_lechei:0.130941)0.796053:0.061597)0.508772:0.039792)0.688596:0.042720,((((((PA6:0.001348,((Galeopterus_variegatus:0.020426,Cynocephalus_volans:0.015253)1.000000:0.171661,((Ptilocercus_lowii:0.075512,Tupaia_glis:0.061774)1.000000:0.053719,(((Tarsius_syrichta:0.144581,(Homo_sapiens:0.155562,Saimiri_sciureus:0.072636)1.000000:0.049677)0.697368:0.023631,(Notharctus_tenebrosus:0.132028,Lemur_catta:0.062883)0.839912:0.039902)1.000000:0.072363,PA26:0.001821)0.699561:0.017463)0.949561:0.032274)1.000000:0.077413)0.916667:0.032019,((Solenodon_paradoxus:0.177599,(((Sorex_araneus:0.139519,Talpa_europaea:0.135911)0.583333:0.046565,Amblysomus_hottentotus:0.247191)0.752193:0.056742,(Erinaceus_europaeus:0.117501,Echinops_telfairi:0.157802)0.697368:0.051929)0.526316:0.030347)0.436404:0.037690,(((((Vulpavus_ovatus:0.018326,Vulpavus_profectus:0.025330)1.000000:0.055636,Sinopa_rapax:0.106935)0.401316:0.040040,(Felis_silvestris:0.088415,Canis_lupus:0.073648)0.995614:0.044927)1.000000:0.073404,(Manis_pentadactyla:0.192716,Metacheiromys_marshi:0.126709)0.995614:0.105572)0.186404:0.032507,(PA7:0.011648,(Pteropus_giganteus:0.177588,(((Myotis_lucifugus:0.062272,Pteronotus_parnellii:0.074898)0.736842:0.026031,(Nycteris_thebaica:0.095225,(Saccopteryx_bilineata:0.052437,Rhinopoma_hardwickii:0.051034)0.741228:0.023501)0.997807:0.031440)0.995614:0.039042,(Icaronycteris_index:0.025547,Onychonycteris_finneyi:0.031712)1.000000:0.082202)0.973684:0.041082)1.000000:0.145686)0.478070:0.019581)0.107456:0.029691)0.074561:0.028592)0.074561:0.027763,((((((Rhombomylus_turpanensis:0.077488,(Gomphos_elkema:0.078684,(Dawsonolagus_antiquus:0.025370,Oryctolagus_cuniculus:0.099254)0.971491:0.033167)1.000000:0.070948)0.614035:0.021324,Tribosphenomys_minutus:0.115408)0.864035:0.033878,((Cocomys_lingchaensis:0.075018,Paramys_delicatus:0.051299)0.381579:0.028427,(((Cavia_porcellus:0.091466,Castor_canadensis:0.084467)0.960526:0.046855,Ictidomys_tridecemlineatus:0.067215)0.473684:0.043992,Rattus_norvegicus:0.149044)0.285088:0.023869)0.640351:0.030848)1.000000:0.091534,((Mesonyx_obtusidens:0.173127,(Rodhocetus_balochistanensis:0.142275,((Mesohippus_bairdi:0.153024,Equus_caballus:0.143150)0.978070:0.060658,((Bos_taurus:0.126143,Lama_glama:0.119683)0.997807:0.056819,((((Artiocetus_clavis:0.134553,(Basilosaurus_cetoides:0.135627,(Tursiops_truncatus:0.169427,Caperea_marginata:0.159584)0.997807:0.109067)0.587719:0.047214)0.953947:0.106811,Archaeotherium_mortoni:0.110613)0.182018:0.036409,Hippopotamus_amphibius:0.173890)0.151316:0.042385,Sus_scrofa:0.175340)0.076754:0.030762)0.067982:0.029494)0.109649:0.026628)0.230263:0.018105)0.899123:0.042877,((Thomashuxleya_externa_comb:0.128769,Carodnia_vieirai:0.118730)0.258772:0.054931,((Didolodus_multicuspis:0.071575,Protolipterna_ellipsodontoides:0.056049)0.607456:0.033409,(((Protungulatum_donnae:0.135463,Apheliscus_insidiosus:0.105570)0.173246:0.031788,Hyopsodus_paulus:0.085136)0.096491:0.027740,Phenacodus_intermedius:0.095624)0.052632:0.015555)0.620614:0.036426)0.122807:0.042184)0.127193:0.040984)0.100877:0.038697,(Leptictis_dakotensis:0.215564,(((((Tamandua_tetradactyla:0.119541,Hapalops_elongatus:0.184206)0.997807:0.084531,(Dasypus_novemcinctus:0.000001,PA39:0.000001)1.000000:0.164175)0.844298:0.086368,PA38:0.000001)0.412281:0.022090,((((Loxodonta_africana:0.183925,(Moeritherium_lyonsi:0.152314,Trichechus_manatus:0.162475)0.331140:0.046506)0.666667:0.087298,Procavia_capensis:0.186249)0.190789:0.044973,PA40:0.002958)0.190789:0.018928,Orycteropus_afer:0.224532)0.078947:0.015202)0.081140:0.038446,Rhynchocyon_cirnei:0.128737)0.083333:0.049256)0.015351:0.028002)0.063596:0.033530,PA5:0.007348)0.061404:0.018503)0.642544:0.042160,PA3:0.005705)0.646930:0.042187,((Ornithorhynchus_anatinus:0.082090,Tachyglossus_aculeatus:0.085346)1.000000:0.137577,PA1:0.000001)0.526316:0.054164)0.460526:0.041337,((Pucadelphys_andinus:0.125371,Didelphis_virginiana:0.041501)1.000000:0.055510,(Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.064388)1.000000:0.064544)0.473684:0.066711)0.997807:0.079419,(Zhangheotherium_quinquecuspidens:0.060125,Henkelotherium_guimarotae:0.067222)0.932018:0.043742)1.000000:0.115673,Haldanodon_exspectatus:0.091547)1.000000:0.081688,Morganucodon_watsoni:0.008518,Morganucodon_oehleri:0.007784);

["skeletal only" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: nodal values = "transfer bootstrap expectation" values]
(((((Eomaia_scansoria:0.050666,(Maelestes_gobiensis:0.094692,(Ukhaatherium_nessovi:0.098999,Zalambdalestes_lechei:0.130941)0.796053:0.061597)0.753289:0.039792)0.888158:0.042720,((((((PA6:0.001348,((Galeopterus_variegatus:0.020426,Cynocephalus_volans:0.015253)1.000000:0.171661,((Ptilocercus_lowii:0.075512,Tupaia_glis:0.061774)1.000000:0.053719,(((Tarsius_syrichta:0.144581,(Homo_sapiens:0.155562,Saimiri_sciureus:0.072636)1.000000:0.049677)0.848684:0.023631,(Notharctus_tenebrosus:0.132028,Lemur_catta:0.062883)0.839912:0.039902)1.000000:0.072363,PA26:0.001821)0.939912:0.017463)0.985589:0.032274)1.000000:0.077413)0.991667:0.032019,((Solenodon_paradoxus:0.177599,(((Sorex_araneus:0.139519,Talpa_europaea:0.135911)0.583333:0.046565,Amblysomus_hottentotus:0.247191)0.876096:0.056742,(Erinaceus_europaeus:0.117501,Echinops_telfairi:0.157802)0.697368:0.051929)0.769189:0.030347)0.781140:0.037690,(((((Vulpavus_ovatus:0.018326,Vulpavus_profectus:0.025330)1.000000:0.055636,Sinopa_rapax:0.106935)0.700658:0.040040,(Felis_silvestris:0.088415,Canis_lupus:0.073648)0.995614:0.044927)1.000000:0.073404,(Manis_pentadactyla:0.192716,Metacheiromys_marshi:0.126709)0.995614:0.105572)0.730263:0.032507,(PA7:0.011648,(Pteropus_giganteus:0.177588,(((Myotis_lucifugus:0.062272,Pteronotus_parnellii:0.074898)0.736842:0.026031,(Nycteris_thebaica:0.095225,(Saccopteryx_bilineata:0.052437,Rhinopoma_hardwickii:0.051034)0.741228:0.023501)0.998904:0.031440)0.997807:0.039042,(Icaronycteris_index:0.025547,Onychonycteris_finneyi:0.031712)1.000000:0.082202)0.995614:0.041082)1.000000:0.145686)0.934759:0.019581)0.713304:0.029691)0.763367:0.028592)0.805099:0.027763,((((((Rhombomylus_turpanensis:0.077488,(Gomphos_elkema:0.078684,(Dawsonolagus_antiquus:0.025370,Oryctolagus_cuniculus:0.099254)0.971491:0.033167)1.000000:0.070948)0.871345:0.021324,Tribosphenomys_minutus:0.115408)0.959978:0.033878,((Cocomys_lingchaensis:0.075018,Paramys_delicatus:0.051299)0.381579:0.028427,(((Cavia_porcellus:0.091466,Castor_canadensis:0.084467)0.960526:0.046855,Ictidomys_tridecemlineatus:0.067215)0.736842:0.043992,Rattus_norvegicus:0.149044)0.647661:0.023869)0.856579:0.030848)1.000000:0.091534,((Mesonyx_obtusidens:0.173127,(Rodhocetus_balochistanensis:0.142275,((Mesohippus_bairdi:0.153024,Equus_caballus:0.143150)0.978070:0.060658,((Bos_taurus:0.126143,Lama_glama:0.119683)0.997807:0.056819,((((Artiocetus_clavis:0.134553,(Basilosaurus_cetoides:0.135627,(Tursiops_truncatus:0.169427,Caperea_marginata:0.159584)0.997807:0.109067)0.793860:0.047214)0.981725:0.106811,Archaeotherium_mortoni:0.110613)0.782895:0.036409,Hippopotamus_amphibius:0.173890)0.700439:0.042385,Sus_scrofa:0.175340)0.623904:0.030762)0.615680:0.029494)0.821491:0.026628)0.900120:0.018105)0.970577:0.042877,((Thomashuxleya_externa_comb:0.128769,Carodnia_vieirai:0.118730)0.258772:0.054931,((Didolodus_multicuspis:0.071575,Protolipterna_ellipsodontoides:0.056049)0.607456:0.033409,(((Protungulatum_donnae:0.135463,Apheliscus_insidiosus:0.105570)0.173246:0.031788,Hyopsodus_paulus:0.085136)0.259868:0.027740,Phenacodus_intermedius:0.095624)0.418860:0.015555)0.903947:0.036426)0.751566:0.042184)0.814035:0.040984)0.817063:0.038697,(Leptictis_dakotensis:0.215564,(((((Tamandua_tetradactyla:0.119541,Hapalops_elongatus:0.184206)0.997807:0.084531,(Dasypus_novemcinctus:0.000001,PA39:0.000001)1.000000:0.164175)0.896199:0.086368,PA38:0.000001)0.799890:0.022090,((((Loxodonta_africana:0.183925,(Moeritherium_lyonsi:0.152314,Trichechus_manatus:0.162475)0.331140:0.046506)0.828947:0.087298,Procavia_capensis:0.186249)0.663012:0.044973,PA40:0.002958)0.582237:0.018928,Orycteropus_afer:0.224532)0.504386:0.015202)0.629386:0.038446,Rhynchocyon_cirnei:0.128737)0.706140:0.049256)0.652778:0.028002)0.851276:0.033530,PA5:0.007348)0.849513:0.018503)0.914611:0.042160,PA3:0.005705)0.909942:0.042187,((Ornithorhynchus_anatinus:0.082090,Tachyglossus_aculeatus:0.085346)1.000000:0.137577,PA1:0.000001)0.763158:0.054164)0.763523:0.041337,((Pucadelphys_andinus:0.125371,Didelphis_virginiana:0.041501)1.000000:0.055510,(Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.064388)1.000000:0.064544)0.746711:0.066711)0.998904:0.079419,(Zhangheotherium_quinquecuspidens:0.060125,Henkelotherium_guimarotae:0.067222)0.932018:0.043742)1.000000:0.115673,Haldanodon_exspectatus:0.091547)1.000000:0.081688,Morganucodon_watsoni:0.008518,Morganucodon_oehleri:0.007784);

["craniodental only" PAs corrected for the O'Leary et al. character ontology: nodal values = standard bootstrap values]
((Haldanodon_exspectatus:0.099527,(((((PA1:0.025687,(PA3:0.003805,((((((((Erinaceus_europaeus:0.099710,(PA24:0.000001,((PA25:0.000001,Sorex_araneus:0.000001)1.000000:0.115640,Talpa_europaea:0.137135)0.880000:0.041513)0.880000:0.046917)0.793333:0.048139,PA23:0.000001)0.786667:0.046064,Solenodon_paradoxus:0.120952)1.000000:0.092868,PA7:0.001681)0.576667:0.017994,(Pteropus_giganteus:0.168475,(((Rhinopoma_hardwickii:0.000001,PA20:0.000001)1.000000:0.049932,((((PA19:0.000001,Saccopteryx_bilineata:0.000001)1.000000:0.045310,(PA18:0.007675,Nycteris_thebaica:0.088614)0.970000:0.014182)0.976667:0.024095,((PA17:0.000001,Myotis_lucifugus:0.050256)0.943333:0.009845,Pteronotus_parnellii:0.074056)0.866667:0.031100)0.543333:0.021787,PA16:0.000001)0.490000:0.013095)0.956667:0.050914,(Onychonycteris_finneyi:0.031379,Icaronycteris_index:0.024967)1.000000:0.099910)0.776667:0.026521)1.000000:0.146754)0.283333:0.021198,(((((PA10:0.020246,(Lama_glama:0.100225,((((Hippopotamus_amphibius:0.000001,PA14:0.000001)1.000000:0.205423,(Sus_scrofa:0.153134,PA12:0.000001)0.590000:0.010247)0.260000:0.034285,(PA13:0.012904,Bos_taurus:0.038859)0.983333:0.064181)0.476667:0.044784,PA11:0.003787)0.363333:0.025858)0.443333:0.026888)0.460000:0.041348,(PA9:0.013762,(Mesohippus_bairdi:0.195209,Equus_caballus:0.106539)0.710000:0.058914)0.316667:0.015141)0.493333:0.046287,(((Rodhocetus_balochistanensis:0.101125,(Artiocetus_clavis:0.125164,(((PA15:0.000001,Tursiops_truncatus:0.000001)1.000000:0.151517,Caperea_marginata:0.149272)1.000000:0.122572,Basilosaurus_cetoides:0.137142)0.590000:0.045713)0.910000:0.047907)0.530000:0.048991,Archaeotherium_mortoni:0.137531)0.183333:0.034103,Mesonyx_obtusidens:0.146324)0.573333:0.078304)0.890000:0.070301,PA8:0.009712)0.450000:0.041794,((Sinopa_rapax:0.106266,(Vulpavus_profectus:0.027119,Vulpavus_ovatus:0.016385)1.000000:0.054650)0.796667:0.053061,((PA21:0.000001,(PA22:0.000001,Felis_silvestris:0.000001)1.000000:0.017621)1.000000:0.058123,Canis_lupus:0.061553)1.000000:0.048607)1.000000:0.071440)0.286667:0.046278)0.376667:0.038264,(PA6:0.000505,((Ptilocercus_lowii:0.070878,(PA27:0.007636,(Tupaia_glis:0.000001,PA32:0.000001)1.000000:0.040255)1.000000:0.025694)1.000000:0.053171,(((Galeopterus_variegatus:0.018319,(PA34:0.000001,Cynocephalus_volans:0.000001)1.000000:0.017128)1.000000:0.143343,(((Notharctus_tenebrosus:0.146345,Lemur_catta:0.049534)0.813333:0.025100,((Tarsius_syrichta:0.125822,(Homo_sapiens:0.153912,(PA36:0.002564,(Saimiri_sciureus:0.000001,PA37:0.000001)1.000000:0.041875)0.910000:0.019850)1.000000:0.050437)0.730000:0.014229,PA35:0.012660)0.830000:0.033170)0.913333:0.029846,PA33:0.000001)1.000000:0.058373)0.970000:0.056237,PA26:0.000001)0.613333:0.028101)1.000000:0.091148)0.806667:0.027819)0.300000:0.029082,((((((((PA28:0.019746,((PA29:0.004072,(Castor_canadensis:0.000001,PA30:0.000001)1.000000:0.073806)0.963333:0.024984,Cavia_porcellus:0.087432)0.570000:0.013806)0.633333:0.022582,(Ictidomys_tridecemlineatus:0.000001,PA31:0.000001)1.000000:0.066575)0.823333:0.047862,Rattus_norvegicus:0.136609)0.940000:0.035617,(Cocomys_lingchaensis:0.071272,Paramys_delicatus:0.053283)0.706667:0.028279)0.696667:0.025938,(Tribosphenomys_minutus:0.109517,(((Dawsonolagus_antiquus:0.025242,Oryctolagus_cuniculus:0.097298)0.986667:0.034908,Gomphos_elkema:0.077826)1.000000:0.063021,Rhombomylus_turpanensis:0.085317)0.686667:0.023166)0.810000:0.032682)1.000000:0.097896,(Leptictis_dakotensis:0.181424,(((Protungulatum_donnae:0.134528,Apheliscus_insidiosus:0.100113)0.183333:0.034882,((Phenacodus_intermedius:0.091019,Hyopsodus_paulus:0.099745)0.436667:0.027339,(Protolipterna_ellipsodontoides:0.049498,Didolodus_multicuspis:0.074945)0.623333:0.031846)0.133333:0.012397)0.500000:0.034419,((Thomashuxleya_externa_comb:0.129070,Carodnia_vieirai:0.107042)0.526667:0.036827,Moeritherium_lyonsi:0.203663)0.313333:0.054832)0.116667:0.029518)0.096667:0.039625)0.090000:0.033829,(((Rhynchocyon_cirnei:0.107851,(((Amblysomus_hottentotus:0.227484,(Echinops_telfairi:0.000001,PA45:0.000001)1.000000:0.116931)0.603333:0.028395,PA44:0.010796)0.973333:0.027827,PA43:0.009669)0.973333:0.027326)0.993333:0.051278,((Orycteropus_afer:0.229563,(((Loxodonta_africana:0.216544,(Trichechus_manatus:0.222191,PA41:0.000001)0.226667:0.000001)0.266667:0.047906,(PA42:0.000001,Procavia_capensis:0.000001)1.000000:0.095676)0.423333:0.128826,PA40:0.002653)0.290000:0.015110)0.176667:0.012870,PA38:0.011748)0.113333:0.027410)0.116667:0.038440,(((Hapalops_elongatus:0.161390,Tamandua_tetradactyla:0.137155)0.996667:0.071258,(PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.175748)0.873333:0.051385,(Metacheiromys_marshi:0.114935,Manis_pentadactyla:0.192287)1.000000:0.073828)0.863333:0.061762)0.233333:0.036416)0.250000:0.031536,PA5:0.001996)0.236667:0.018291)0.810000:0.042144)0.823333:0.027626)0.700000:0.043229,((PA4:0.000001,Dromiciops_gliroides:0.000001)1.000000:0.071936,(Pucadelphys_andinus:0.119618,Didelphis_virginiana:0.046937)0.996667:0.056321)1.000000:0.065168)0.700000:0.052153,(((Zalambdalestes_lechei:0.130192,Ukhaatherium_nessovi:0.096489)0.950000:0.061596,Maelestes_gobiensis:0.094076)0.583333:0.039979,Eomaia_scansoria:0.052404)0.610000:0.044890)0.920000:0.079014,(Zhangheotherium_quinquecuspidens:0.059413,Henkelotherium_guimarotae:0.067581)0.883333:0.040825)0.820000:0.057562,(Tachyglossus_aculeatus:0.084563,(Ornithorhynchus_anatinus:0.000001,PA2:0.000001)1.000000:0.076522)1.000000:0.131241)1.000000:0.070475)1.000000:0.077261,Morganucodon_watsoni:0.008070,Morganucodon_oehleri:0.007934);

["craniodental only" PAs corrected for the O'Leary et al. character ontology: nodal values = "transfer bootstrap expectation" values]
((Haldanodon_exspectatus:0.099527,(((((PA1:0.025687,(PA3:0.003805,((((((((Erinaceus_europaeus:0.099710,(PA24:0.000001,((PA25:0.000001,Sorex_araneus:0.000001)1.000000:0.115640,Talpa_europaea:0.137135)0.940000:0.041513)0.960000:0.046917)0.948333:0.048139,PA23:0.000001)0.957333:0.046064,Solenodon_paradoxus:0.120952)1.000000:0.092868,PA7:0.001681)0.939524:0.017994,(Pteropus_giganteus:0.168475,(((Rhinopoma_hardwickii:0.000001,PA20:0.000001)1.000000:0.049932,((((PA19:0.000001,Saccopteryx_bilineata:0.000001)1.000000:0.045310,(PA18:0.007675,Nycteris_thebaica:0.088614)0.970000:0.014182)0.985556:0.024095,((PA17:0.000001,Myotis_lucifugus:0.050256)0.943333:0.009845,Pteronotus_parnellii:0.074056)0.931667:0.031100)0.778889:0.021787,PA16:0.000001)0.861905:0.013095)0.990370:0.050914,(Onychonycteris_finneyi:0.031379,Icaronycteris_index:0.024967)1.000000:0.099910)0.979697:0.026521)1.000000:0.146754)0.719000:0.021198,(((((PA10:0.020246,(Lama_glama:0.100225,((((Hippopotamus_amphibius:0.000001,PA14:0.000001)1.000000:0.205423,(Sus_scrofa:0.153134,PA12:0.000001)0.590000:0.010247)0.512222:0.034285,(PA13:0.012904,Bos_taurus:0.038859)0.983333:0.064181)0.640000:0.044784,PA11:0.003787)0.616667:0.025858)0.688095:0.026888)0.723750:0.041348,(PA9:0.013762,(Mesohippus_bairdi:0.195209,Equus_caballus:0.106539)0.710000:0.058914)0.546667:0.015141)0.791818:0.046287,(((Rodhocetus_balochistanensis:0.101125,(Artiocetus_clavis:0.125164,(((PA15:0.000001,Tursiops_truncatus:0.000001)1.000000:0.151517,Caperea_marginata:0.149272)1.000000:0.122572,Basilosaurus_cetoides:0.137142)0.863333:0.045713)0.971667:0.047907)0.893333:0.048991,Archaeotherium_mortoni:0.137531)0.832222:0.034103,Mesonyx_obtusidens:0.146324)0.892381:0.078304)0.966842:0.070301,PA8:0.009712)0.944167:0.041794,((Sinopa_rapax:0.106266,(Vulpavus_profectus:0.027119,Vulpavus_ovatus:0.016385)1.000000:0.054650)0.898333:0.053061,((PA21:0.000001,(PA22:0.000001,Felis_silvestris:0.000001)1.000000:0.017621)1.000000:0.058123,Canis_lupus:0.061553)1.000000:0.048607)1.000000:0.071440)0.786296:0.046278)0.822847:0.038264,(PA6:0.000505,((Ptilocercus_lowii:0.070878,(PA27:0.007636,(Tupaia_glis:0.000001,PA32:0.000001)1.000000:0.040255)1.000000:0.025694)1.000000:0.053171,(((Galeopterus_variegatus:0.018319,(PA34:0.000001,Cynocephalus_volans:0.000001)1.000000:0.017128)1.000000:0.143343,(((Notharctus_tenebrosus:0.146345,Lemur_catta:0.049534)0.813333:0.025100,((Tarsius_syrichta:0.125822,(Homo_sapiens:0.153912,(PA36:0.002564,(Saimiri_sciureus:0.000001,PA37:0.000001)1.000000:0.041875)0.955000:0.019850)1.000000:0.050437)0.932500:0.014229,PA35:0.012660)0.933333:0.033170)0.987619:0.029846,PA33:0.000001)1.000000:0.058373)0.991818:0.056237,PA26:0.000001)0.960556:0.028101)1.000000:0.091148)0.988627:0.027819)0.807672:0.029082,((((((((PA28:0.019746,((PA29:0.004072,(Castor_canadensis:0.000001,PA30:0.000001)1.000000:0.073806)0.981667:0.024984,Cavia_porcellus:0.087432)0.844444:0.013806)0.820000:0.022582,(Ictidomys_tridecemlineatus:0.000001,PA31:0.000001)1.000000:0.066575)0.970556:0.047862,Rattus_norvegicus:0.136609)0.991429:0.035617,(Cocomys_lingchaensis:0.071272,Paramys_delicatus:0.053283)0.706667:0.028279)0.933704:0.025938,(Tribosphenomys_minutus:0.109517,(((Dawsonolagus_antiquus:0.025242,Oryctolagus_cuniculus:0.097298)0.986667:0.034908,Gomphos_elkema:0.077826)1.000000:0.063021,Rhombomylus_turpanensis:0.085317)0.895556:0.023166)0.951667:0.032682)1.000000:0.097896,(Leptictis_dakotensis:0.181424,(((Protungulatum_donnae:0.134528,Apheliscus_insidiosus:0.100113)0.183333:0.034882,((Phenacodus_intermedius:0.091019,Hyopsodus_paulus:0.099745)0.436667:0.027339,(Protolipterna_ellipsodontoides:0.049498,Didolodus_multicuspis:0.074945)0.623333:0.031846)0.537778:0.012397)0.839333:0.034419,((Thomashuxleya_externa_comb:0.129070,Carodnia_vieirai:0.107042)0.526667:0.036827,Moeritherium_lyonsi:0.203663)0.546667:0.054832)0.755833:0.029518)0.736296:0.039625)0.722917:0.033829,(((Rhynchocyon_cirnei:0.107851,(((Amblysomus_hottentotus:0.227484,(Echinops_telfairi:0.000001,PA45:0.000001)1.000000:0.116931)0.801667:0.028395,PA44:0.010796)0.982222:0.027827,PA43:0.009669)0.991667:0.027326)0.997333:0.051278,((Orycteropus_afer:0.229563,(((Loxodonta_africana:0.216544,(Trichechus_manatus:0.222191,PA41:0.000001)0.226667:0.000001)0.321667:0.047906,(PA42:0.000001,Procavia_capensis:0.000001)1.000000:0.095676)0.815000:0.128826,PA40:0.002653)0.797333:0.015110)0.735556:0.012870,PA38:0.011748)0.680476:0.027410)0.653077:0.038440,(((Hapalops_elongatus:0.161390,Tamandua_tetradactyla:0.137155)0.996667:0.071258,(PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.175748)0.915556:0.051385,(Metacheiromys_marshi:0.114935,Manis_pentadactyla:0.192287)1.000000:0.073828)0.946000:0.061762)0.906491:0.036416)0.743182:0.031536,PA5:0.001996)0.744741:0.018291)0.970980:0.042144)0.970000:0.027626)0.954444:0.043229,((PA4:0.000001,Dromiciops_gliroides:0.000001)1.000000:0.071936,(Pucadelphys_andinus:0.119618,Didelphis_virginiana:0.046937)0.996667:0.056321)1.000000:0.065168)0.914848:0.052153,(((Zalambdalestes_lechei:0.130192,Ukhaatherium_nessovi:0.096489)0.950000:0.061596,Maelestes_gobiensis:0.094076)0.791667:0.039979,Eomaia_scansoria:0.052404)0.860000:0.044890)0.965714:0.079014,(Zhangheotherium_quinquecuspidens:0.059413,Henkelotherium_guimarotae:0.067581)0.883333:0.040825)0.912000:0.057562,(Tachyglossus_aculeatus:0.084563,(Ornithorhynchus_anatinus:0.000001,PA2:0.000001)1.000000:0.076522)1.000000:0.131241)1.000000:0.070475)1.000000:0.077261,Morganucodon_watsoni:0.008070,Morganucodon_oehleri:0.007934);

["craniodental only" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: nodal values = standard bootstrap values]
((Haldanodon_exspectatus:0.099690,((((((Didelphis_virginiana:0.047013,Pucadelphys_andinus:0.119236)1.000000:0.056621,(Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.071536)1.000000:0.065420,(PA1:0.025665,(PA3:0.003707,(((((Leptictis_dakotensis:0.181302,(((Protungulatum_donnae:0.136832,Apheliscus_insidiosus:0.099702)0.197222:0.034426,((Protolipterna_ellipsodontoides:0.050144,Didolodus_multicuspis:0.075086)0.597222:0.032822,(Hyopsodus_paulus:0.099461,Phenacodus_intermedius:0.091320)0.450000:0.026227)0.144444:0.013104)0.561111:0.033075,(Thomashuxleya_externa_comb:0.137215,(Moeritherium_lyonsi:0.184140,Carodnia_vieirai:0.093167)0.272222:0.046393)0.316667:0.061522)0.144444:0.030225)0.083333:0.039373,((Tribosphenomys_minutus:0.110180,((Gomphos_elkema:0.079404,(Dawsonolagus_antiquus:0.025365,Oryctolagus_cuniculus:0.096555)0.961111:0.033815)1.000000:0.062191,Rhombomylus_turpanensis:0.086194)0.680556:0.022358)0.838889:0.031515,(((Ictidomys_tridecemlineatus:0.069903,((Cavia_porcellus:0.087576,Castor_canadensis:0.091030)0.655556:0.012960,PA28:0.000001)0.872222:0.036722)0.725000:0.044644,Rattus_norvegicus:0.137448)0.816667:0.035419,(Paramys_delicatus:0.053064,Cocomys_lingchaensis:0.071050)0.711111:0.029515)0.700000:0.024887)1.000000:0.097901)0.058333:0.032005,((((Echinops_telfairi:0.118073,(Amblysomus_hottentotus:0.243535,PA44:0.000001)0.927778:0.020403)0.922222:0.024929,PA43:0.006048)0.919444:0.031940,Rhynchocyon_cirnei:0.109949)0.980556:0.045745,(((Orycteropus_afer:0.220520,((Metacheiromys_marshi:0.117645,Manis_pentadactyla:0.191625)0.994444:0.080654,((Hapalops_elongatus:0.168277,Tamandua_tetradactyla:0.133141)0.997222:0.072842,(PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.170392)0.888889:0.045764)0.813889:0.042826)0.302778:0.032448,PA38:0.002447)0.275000:0.014908,(((PA42:0.000001,Procavia_capensis:0.000001)1.000000:0.096602,(Loxodonta_africana:0.216365,(Trichechus_manatus:0.220164,PA41:0.000001)0.216667:0.000001)0.255556:0.047958)0.438889:0.130056,PA40:0.003138)0.308333:0.021706)0.152778:0.030061)0.255556:0.063647)0.186111:0.034710,PA5:0.001887)0.163889:0.018275,(((((Sinopa_rapax:0.107315,(Vulpavus_ovatus:0.016436,Vulpavus_profectus:0.026993)1.000000:0.053615)0.830556:0.052561,(Canis_lupus:0.060963,(Felis_silvestris:0.022014,PA21:0.000001)1.000000:0.056891)0.997222:0.048691)1.000000:0.072235,((PA9:0.018286,(((PA10:0.000168,(Bos_taurus:0.136492,Lama_glama:0.113417)0.650000:0.018820)0.963889:0.024126,((Hippopotamus_amphibius:0.160741,(((Rodhocetus_balochistanensis:0.102540,(Basilosaurus_cetoides:0.151944,(Artiocetus_clavis:0.107607,(Tursiops_truncatus:0.161384,Caperea_marginata:0.163696)1.000000:0.104988)0.391667:0.039937)0.963889:0.059935)0.594444:0.038081,Mesonyx_obtusidens:0.146870)0.405556:0.044052,Archaeotherium_mortoni:0.119979)0.483333:0.050814)0.430556:0.056336,Sus_scrofa:0.157788)0.363889:0.038770)0.380556:0.032960,(Mesohippus_bairdi:0.197245,Equus_caballus:0.103767)0.802778:0.049886)0.358333:0.024009)0.980556:0.111887,PA8:0.007238)0.369444:0.039794)0.227778:0.047190,((PA7:0.000658,(((Sorex_araneus:0.130785,Talpa_europaea:0.152496)0.991667:0.071764,Erinaceus_europaeus:0.155629)0.625000:0.037931,Solenodon_paradoxus:0.177150)0.766667:0.035596)0.713889:0.021738,(Pteropus_giganteus:0.179367,((Onychonycteris_finneyi:0.030870,Icaronycteris_index:0.026701)1.000000:0.078289,((Nycteris_thebaica:0.093613,(Saccopteryx_bilineata:0.052045,Rhinopoma_hardwickii:0.051134)0.725000:0.023609)1.000000:0.030376,(Pteronotus_parnellii:0.075175,Myotis_lucifugus:0.061828)0.800000:0.027209)0.997222:0.040145)0.988889:0.038652)1.000000:0.138403)0.302778:0.021460)0.252778:0.037075,(((Ptilocercus_lowii:0.071220,(PA27:0.005874,Tupaia_glis:0.040493)1.000000:0.026324)1.000000:0.053982,(PA26:0.000001,((Galeopterus_variegatus:0.018141,Cynocephalus_volans:0.017394)1.000000:0.143840,((((Notharctus_tenebrosus:0.114533,Tarsius_syrichta:0.133131)0.691667:0.043701,Lemur_catta:0.069149)0.602778:0.028442,PA33:0.000001)0.400000:0.004141,(Saimiri_sciureus:0.069127,Homo_sapiens:0.156755)1.000000:0.055120)1.000000:0.061198)0.966667:0.054117)0.600000:0.028056)1.000000:0.090847,PA6:0.000569)0.802778:0.028015)0.197222:0.028593)0.827778:0.041825)0.838889:0.027461)0.700000:0.042850)0.700000:0.051396,((Maelestes_gobiensis:0.094399,(Zalambdalestes_lechei:0.129879,Ukhaatherium_nessovi:0.096431)0.952778:0.061396)0.647222:0.039741,Eomaia_scansoria:0.052885)0.600000:0.044727)0.919444:0.078907,(Henkelotherium_guimarotae:0.067609,Zhangheotherium_quinquecuspidens:0.059333)0.891667:0.040997)0.830556:0.057549,(Tachyglossus_aculeatus:0.087542,Ornithorhynchus_anatinus:0.079114)1.000000:0.128215)1.000000:0.070884)1.000000:0.077211,Morganucodon_watsoni:0.008215,Morganucodon_oehleri:0.007782);

["craniodental only" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: nodal values = "transfer bootstrap expectation" values]
((Haldanodon_exspectatus:0.099690,((((((Didelphis_virginiana:0.047013,Pucadelphys_andinus:0.119236)1.000000:0.056621,(Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.071536)1.000000:0.065420,(PA1:0.025665,(PA3:0.003707,(((((Leptictis_dakotensis:0.181302,(((Protungulatum_donnae:0.136832,Apheliscus_insidiosus:0.099702)0.197222:0.034426,((Protolipterna_ellipsodontoides:0.050144,Didolodus_multicuspis:0.075086)0.597222:0.032822,(Hyopsodus_paulus:0.099461,Phenacodus_intermedius:0.091320)0.450000:0.026227)0.546296:0.013104)0.869444:0.033075,(Thomashuxleya_externa_comb:0.137215,(Moeritherium_lyonsi:0.184140,Carodnia_vieirai:0.093167)0.272222:0.046393)0.547222:0.061522)0.774306:0.030225)0.749691:0.039373,((Tribosphenomys_minutus:0.110180,((Gomphos_elkema:0.079404,(Dawsonolagus_antiquus:0.025365,Oryctolagus_cuniculus:0.096555)0.961111:0.033815)1.000000:0.062191,Rhombomylus_turpanensis:0.086194)0.893519:0.022358)0.958333:0.031515,(((Ictidomys_tridecemlineatus:0.069903,((Cavia_porcellus:0.087576,Castor_canadensis:0.091030)0.655556:0.012960,PA28:0.000001)0.934722:0.036722)0.908333:0.044644,Rattus_norvegicus:0.137448)0.947222:0.035419,(Paramys_delicatus:0.053064,Cocomys_lingchaensis:0.071050)0.711111:0.029515)0.901389:0.024887)1.000000:0.097901)0.650794:0.032005,((((Echinops_telfairi:0.118073,(Amblysomus_hottentotus:0.243535,PA44:0.000001)0.927778:0.020403)0.956944:0.024929,PA43:0.006048)0.967593:0.031940,Rhynchocyon_cirnei:0.109949)0.991667:0.045745,(((Orycteropus_afer:0.220520,((Metacheiromys_marshi:0.117645,Manis_pentadactyla:0.191625)0.994444:0.080654,((Hapalops_elongatus:0.168277,Tamandua_tetradactyla:0.133141)0.997222:0.072842,(PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.170392)0.925926:0.045764)0.926111:0.042826)0.824074:0.032448,PA38:0.002447)0.800794:0.014908,(((PA42:0.000001,Procavia_capensis:0.000001)1.000000:0.096602,(Loxodonta_africana:0.216365,(Trichechus_manatus:0.220164,PA41:0.000001)0.216667:0.000001)0.308333:0.047958)0.815972:0.130056,PA40:0.003138)0.800000:0.021706)0.712821:0.030061)0.896296:0.063647)0.725069:0.034710,PA5:0.001887)0.727304:0.018275,(((((Sinopa_rapax:0.107315,(Vulpavus_ovatus:0.016436,Vulpavus_profectus:0.026993)1.000000:0.053615)0.915278:0.052561,(Canis_lupus:0.060963,(Felis_silvestris:0.022014,PA21:0.000001)1.000000:0.056891)0.998611:0.048691)1.000000:0.072235,((PA9:0.018286,(((PA10:0.000168,(Bos_taurus:0.136492,Lama_glama:0.113417)0.650000:0.018820)0.981944:0.024126,((Hippopotamus_amphibius:0.160741,(((Rodhocetus_balochistanensis:0.102540,(Basilosaurus_cetoides:0.151944,(Artiocetus_clavis:0.107607,(Tursiops_truncatus:0.161384,Caperea_marginata:0.163696)1.000000:0.104988)0.695833:0.039937)0.987963:0.059935)0.897222:0.038081,Mesonyx_obtusidens:0.146870)0.830000:0.044052,Archaeotherium_mortoni:0.119979)0.848148:0.050814)0.834127:0.056336,Sus_scrofa:0.157788)0.785764:0.038770)0.839646:0.032960,(Mesohippus_bairdi:0.197245,Equus_caballus:0.103767)0.802778:0.049886)0.945726:0.024009)0.994841:0.111887,PA8:0.007238)0.953889:0.039794)0.757540:0.047190,((PA7:0.000658,(((Sorex_araneus:0.130785,Talpa_europaea:0.152496)0.991667:0.071764,Erinaceus_europaeus:0.155629)0.809722:0.037931,Solenodon_paradoxus:0.177150)0.910185:0.035596)0.884722:0.021738,(Pteropus_giganteus:0.179367,((Onychonycteris_finneyi:0.030870,Icaronycteris_index:0.026701)1.000000:0.078289,((Nycteris_thebaica:0.093613,(Saccopteryx_bilineata:0.052045,Rhinopoma_hardwickii:0.051134)0.725000:0.023609)1.000000:0.030376,(Pteronotus_parnellii:0.075175,Myotis_lucifugus:0.061828)0.800000:0.027209)0.998611:0.040145)0.998148:0.038652)1.000000:0.138403)0.738194:0.021460)0.749673:0.037075,(((Ptilocercus_lowii:0.071220,(PA27:0.005874,Tupaia_glis:0.040493)1.000000:0.026324)1.000000:0.053982,(PA26:0.000001,((Galeopterus_variegatus:0.018141,Cynocephalus_volans:0.017394)1.000000:0.143840,((((Notharctus_tenebrosus:0.114533,Tarsius_syrichta:0.133131)0.691667:0.043701,Lemur_catta:0.069149)0.798611:0.028442,PA33:0.000001)0.705556:0.004141,(Saimiri_sciureus:0.069127,Homo_sapiens:0.156755)1.000000:0.055120)1.000000:0.061198)0.990476:0.054117)0.941667:0.028056)1.000000:0.090847,PA6:0.000569)0.983565:0.028015)0.758452:0.028593)0.974132:0.041825)0.972963:0.027461)0.956349:0.042850)0.916389:0.051396,((Maelestes_gobiensis:0.094399,(Zalambdalestes_lechei:0.129879,Ukhaatherium_nessovi:0.096431)0.952778:0.061396)0.822222:0.039741,Eomaia_scansoria:0.052885)0.855556:0.044727)0.973148:0.078907,(Henkelotherium_guimarotae:0.067609,Zhangheotherium_quinquecuspidens:0.059333)0.891667:0.040997)0.915278:0.057549,(Tachyglossus_aculeatus:0.087542,Ornithorhynchus_anatinus:0.079114)1.000000:0.128215)1.000000:0.070884)1.000000:0.077211,Morganucodon_watsoni:0.008215,Morganucodon_oehleri:0.007782);

["craniodental only" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: nodal values = standard bootstrap values]
((((((((PA4:0.000001,Dromiciops_gliroides:0.000001)1.000000:0.071422,(Pucadelphys_andinus:0.119968,Didelphis_virginiana:0.047020)0.997549:0.057652)1.000000:0.064992,(PA1:0.025860,(((((PA7:0.011141,(Pteropus_giganteus:0.180423,((Onychonycteris_finneyi:0.030429,Icaronycteris_index:0.027077)1.000000:0.080535,((Nycteris_thebaica:0.094469,(Saccopteryx_bilineata:0.052405,Rhinopoma_hardwickii:0.051079)0.742647:0.023638)0.997549:0.031043,(Pteronotus_parnellii:0.075609,Myotis_lucifugus:0.061705)0.750000:0.026299)1.000000:0.038561)0.992647:0.040822)1.000000:0.124819)0.725490:0.037908,(Solenodon_paradoxus:0.177568,((Echinops_telfairi:0.154223,Erinaceus_europaeus:0.119411)0.845588:0.052940,((Amblysomus_hottentotus:0.235912,Talpa_europaea:0.131840)0.470588:0.047034,Sorex_araneus:0.145646)0.882353:0.059434)0.625000:0.027646)0.696078:0.030204)0.507353:0.039601,(PA6:0.000939,((Galeopterus_variegatus:0.019970,Cynocephalus_volans:0.015612)1.000000:0.169159,((Tupaia_glis:0.064466,Ptilocercus_lowii:0.073273)1.000000:0.054595,(((Tarsius_syrichta:0.142777,(Homo_sapiens:0.155317,Saimiri_sciureus:0.073518)1.000000:0.051064)0.678922:0.023411,(Notharctus_tenebrosus:0.129957,Lemur_catta:0.065479)0.857843:0.038686)1.000000:0.068205,PA26:0.004094)0.573529:0.021462)0.801471:0.032407)1.000000:0.070629)0.928922:0.033274)0.284314:0.026289,((((Rhynchocyon_cirnei:0.140489,(((Procavia_capensis:0.226711,PA40:0.003770)0.220588:0.015551,Orycteropus_afer:0.223561)0.117647:0.014168,PA38:0.010618)0.100490:0.028021)0.281863:0.047840,((Metacheiromys_marshi:0.116798,Manis_pentadactyla:0.189414)0.990196:0.070177,((Hapalops_elongatus:0.161739,Tamandua_tetradactyla:0.137435)0.992647:0.072690,(PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.175408)0.865196:0.053624)0.843137:0.060471)0.495098:0.038572,(((Leptictis_dakotensis:0.188870,(Protungulatum_donnae:0.132222,(Sinopa_rapax:0.088509,((Canis_lupus:0.065577,Felis_silvestris:0.097874)0.997549:0.053543,(Vulpavus_profectus:0.025976,Vulpavus_ovatus:0.017792)1.000000:0.059971)0.725490:0.026397)1.000000:0.062315)0.117647:0.019109)0.115196:0.031664,((((Protolipterna_ellipsodontoides:0.054780,Didolodus_multicuspis:0.073774)0.605392:0.034795,(Hyopsodus_paulus:0.105475,Phenacodus_intermedius:0.086726)0.490196:0.025691)0.296569:0.014859,Apheliscus_insidiosus:0.115610)0.536765:0.028879,(((Carodnia_vieirai:0.084317,(Moeritherium_lyonsi:0.139182,(Loxodonta_africana:0.173882,Trichechus_manatus:0.191396)0.406863:0.035702)0.691176:0.042657)0.845588:0.051718,Thomashuxleya_externa_comb:0.153004)0.529412:0.038778,(((((Artiocetus_clavis:0.122647,((Caperea_marginata:0.160215,Tursiops_truncatus:0.168676)1.000000:0.114116,Basilosaurus_cetoides:0.141166)0.553922:0.045226)0.941176:0.094770,Archaeotherium_mortoni:0.119498)0.151961:0.031970,Mesonyx_obtusidens:0.149848)0.075980:0.042261,((Mesohippus_bairdi:0.150474,Equus_caballus:0.145053)0.968137:0.063617,((Lama_glama:0.119936,Bos_taurus:0.126100)0.995098:0.059239,(Hippopotamus_amphibius:0.190718,Sus_scrofa:0.160607)0.208333:0.037749)0.095588:0.028459)0.284314:0.043451)0.056373:0.010712,Rodhocetus_balochistanensis:0.149589)0.889706:0.036941)0.588235:0.041922)0.183824:0.035450)0.083333:0.028589,((Tribosphenomys_minutus:0.107899,(Rhombomylus_turpanensis:0.079882,((Oryctolagus_cuniculus:0.099017,Dawsonolagus_antiquus:0.025321)0.970588:0.035300,Gomphos_elkema:0.076290)1.000000:0.069053)0.634804:0.023954)0.855392:0.035139,((Cocomys_lingchaensis:0.073250,Paramys_delicatus:0.052938)0.389706:0.023162,(Rattus_norvegicus:0.151249,(Ictidomys_tridecemlineatus:0.064489,(Castor_canadensis:0.084936,Cavia_porcellus:0.090668)0.948529:0.048742)0.507353:0.045570)0.299020:0.023385)0.639706:0.031751)1.000000:0.094829)0.156863:0.033623)0.240196:0.027749,PA5:0.005019)0.242647:0.020855)0.752451:0.038331,PA3:0.003839)0.789216:0.027512)0.700980:0.043269)0.700980:0.051587,((Maelestes_gobiensis:0.093433,(Zalambdalestes_lechei:0.130551,Ukhaatherium_nessovi:0.097076)0.955882:0.062161)0.627451:0.040143,Eomaia_scansoria:0.051848)0.566176:0.043846)0.928922:0.080130,(Zhangheotherium_quinquecuspidens:0.058990,Henkelotherium_guimarotae:0.068172)0.887255:0.040805)0.860294:0.057448,(Tachyglossus_aculeatus:0.087992,Ornithorhynchus_anatinus:0.079160)1.000000:0.128594)1.000000:0.070380,Haldanodon_exspectatus:0.099716)1.000000:0.077574,Morganucodon_watsoni:0.008229,Morganucodon_oehleri:0.007842);

["craniodental only" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: nodal values = "transfer bootstrap expectation" values]
((((((((PA4:0.000001,Dromiciops_gliroides:0.000001)1.000000:0.071422,(Pucadelphys_andinus:0.119968,Didelphis_virginiana:0.047020)0.997549:0.057652)1.000000:0.064992,(PA1:0.025860,(((((PA7:0.011141,(Pteropus_giganteus:0.180423,((Onychonycteris_finneyi:0.030429,Icaronycteris_index:0.027077)1.000000:0.080535,((Nycteris_thebaica:0.094469,(Saccopteryx_bilineata:0.052405,Rhinopoma_hardwickii:0.051079)0.742647:0.023638)0.998775:0.031043,(Pteronotus_parnellii:0.075609,Myotis_lucifugus:0.061705)0.750000:0.026299)1.000000:0.038561)0.998775:0.040822)1.000000:0.124819)0.965686:0.037908,(Solenodon_paradoxus:0.177568,((Echinops_telfairi:0.154223,Erinaceus_europaeus:0.119411)0.845588:0.052940,((Amblysomus_hottentotus:0.235912,Talpa_europaea:0.131840)0.470588:0.047034,Sorex_araneus:0.145646)0.941176:0.059434)0.853554:0.027646)0.890686:0.030204)0.832808:0.039601,(PA6:0.000939,((Galeopterus_variegatus:0.019970,Cynocephalus_volans:0.015612)1.000000:0.169159,((Tupaia_glis:0.064466,Ptilocercus_lowii:0.073273)1.000000:0.054595,(((Tarsius_syrichta:0.142777,(Homo_sapiens:0.155317,Saimiri_sciureus:0.073518)1.000000:0.051064)0.839461:0.023411,(Notharctus_tenebrosus:0.129957,Lemur_catta:0.065479)0.857843:0.038686)1.000000:0.068205,PA26:0.004094)0.914706:0.021462)0.943277:0.032407)1.000000:0.070629)0.992892:0.033274)0.756765:0.026289,((((Rhynchocyon_cirnei:0.140489,(((Procavia_capensis:0.226711,PA40:0.003770)0.220588:0.015551,Orycteropus_afer:0.223561)0.324755:0.014168,PA38:0.010618)0.369281:0.028021)0.454044:0.047840,((Metacheiromys_marshi:0.116798,Manis_pentadactyla:0.189414)0.990196:0.070177,((Hapalops_elongatus:0.161739,Tamandua_tetradactyla:0.137435)0.992647:0.072690,(PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.175408)0.910131:0.053624)0.940196:0.060471)0.847304:0.038572,(((Leptictis_dakotensis:0.188870,(Protungulatum_donnae:0.132222,(Sinopa_rapax:0.088509,((Canis_lupus:0.065577,Felis_silvestris:0.097874)0.997549:0.053543,(Vulpavus_profectus:0.025976,Vulpavus_ovatus:0.017792)1.000000:0.059971)0.908497:0.026397)1.000000:0.062315)0.823529:0.019109)0.732026:0.031664,((((Protolipterna_ellipsodontoides:0.054780,Didolodus_multicuspis:0.073774)0.605392:0.034795,(Hyopsodus_paulus:0.105475,Phenacodus_intermedius:0.086726)0.490196:0.025691)0.671569:0.014859,Apheliscus_insidiosus:0.115610)0.869485:0.028879,(((Carodnia_vieirai:0.084317,(Moeritherium_lyonsi:0.139182,(Loxodonta_africana:0.173882,Trichechus_manatus:0.191396)0.406863:0.035702)0.840686:0.042657)0.942810:0.051718,Thomashuxleya_externa_comb:0.153004)0.859681:0.038778,(((((Artiocetus_clavis:0.122647,((Caperea_marginata:0.160215,Tursiops_truncatus:0.168676)1.000000:0.114116,Basilosaurus_cetoides:0.141166)0.776961:0.045226)0.975490:0.094770,Archaeotherium_mortoni:0.119498)0.771446:0.031970,Mesonyx_obtusidens:0.149848)0.686765:0.042261,((Mesohippus_bairdi:0.150474,Equus_caballus:0.145053)0.968137:0.063617,((Lama_glama:0.119936,Bos_taurus:0.126100)0.995098:0.059239,(Hippopotamus_amphibius:0.190718,Sus_scrofa:0.160607)0.208333:0.037749)0.477124:0.028459)0.690686:0.043451)0.883913:0.010712,Rodhocetus_balochistanensis:0.149589)0.969567:0.036941)0.918397:0.041922)0.854724:0.035450)0.735463:0.028589,((Tribosphenomys_minutus:0.107899,(Rhombomylus_turpanensis:0.079882,((Oryctolagus_cuniculus:0.099017,Dawsonolagus_antiquus:0.025321)0.970588:0.035300,Gomphos_elkema:0.076290)1.000000:0.069053)0.878268:0.023954)0.960784:0.035139,((Cocomys_lingchaensis:0.073250,Paramys_delicatus:0.052938)0.389706:0.023162,(Rattus_norvegicus:0.151249,(Ictidomys_tridecemlineatus:0.064489,(Castor_canadensis:0.084936,Cavia_porcellus:0.090668)0.948529:0.048742)0.753676:0.045570)0.665850:0.023385)0.860294:0.031751)1.000000:0.094829)0.825980:0.033623)0.854081:0.027749,PA5:0.005019)0.849090:0.020855)0.958946:0.038331,PA3:0.003839)0.959150:0.027512)0.943627:0.043269)0.911275:0.051587,((Maelestes_gobiensis:0.093433,(Zalambdalestes_lechei:0.130551,Ukhaatherium_nessovi:0.097076)0.955882:0.062161)0.813725:0.040143,Eomaia_scansoria:0.051848)0.843954:0.043846)0.976307:0.080130,(Zhangheotherium_quinquecuspidens:0.058990,Henkelotherium_guimarotae:0.068172)0.887255:0.040805)0.930147:0.057448,(Tachyglossus_aculeatus:0.087992,Ornithorhynchus_anatinus:0.079160)1.000000:0.128594)1.000000:0.070380,Haldanodon_exspectatus:0.099716)1.000000:0.077574,Morganucodon_watsoni:0.008229,Morganucodon_oehleri:0.007842);

["dental only" PAs corrected for the O'Leary et al. character ontology: nodal values = standard bootstrap values]
((((Tachyglossus_aculeatus:0.089091,(PA2:0.000001,Ornithorhynchus_anatinus:0.000001)1.000000:0.079871)1.000000:0.136108,(((((Pucadelphys_andinus:0.115891,Didelphis_virginiana:0.052945)0.980556:0.057909,(Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.076726)1.000000:0.066456,((((((((((PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.184910,(Hapalops_elongatus:0.160648,Tamandua_tetradactyla:0.144268)1.000000:0.078499)0.983333:0.057124,(Metacheiromys_marshi:0.117476,Manis_pentadactyla:0.192029)0.961111:0.061191)0.872222:0.044126,Orycteropus_afer:0.232959)0.833333:0.058461,((((Rattus_norvegicus:0.146435,((PA31:0.000001,Ictidomys_tridecemlineatus:0.000001)1.000000:0.060622,((Cavia_porcellus:0.070340,PA28:0.005067)0.616667:0.025921,(PA29:0.003594,(PA30:0.000001,Castor_canadensis:0.000001)1.000000:0.076848)0.902778:0.018578)0.880556:0.034645)0.866667:0.051210)0.725000:0.028559,(Cocomys_lingchaensis:0.073284,Paramys_delicatus:0.052833)0.544444:0.030694)0.758333:0.031304,((Tribosphenomys_minutus:0.119274,Rhombomylus_turpanensis:0.066816)0.236111:0.022397,((Dawsonolagus_antiquus:0.025436,Oryctolagus_cuniculus:0.100239)0.975000:0.033817,Gomphos_elkema:0.080236)1.000000:0.069179)0.836111:0.046631)1.000000:0.090575,((Thomashuxleya_externa_comb:0.156600,((Mesonyx_obtusidens:0.173312,(((((Caperea_marginata:0.171521,(Tursiops_truncatus:0.000001,PA15:0.000001)0.980556:0.169959)0.980556:0.113045,Basilosaurus_cetoides:0.147998)0.602778:0.053288,Artiocetus_clavis:0.124237)0.922222:0.062851,Rodhocetus_balochistanensis:0.103411)0.469444:0.044750,(Archaeotherium_mortoni:0.140562,((PA14:0.000001,Hippopotamus_amphibius:0.000001)1.000000:0.177719,(((PA10:0.013937,((Mesohippus_bairdi:0.191032,Equus_caballus:0.105936)0.572222:0.033170,PA9:0.029627)0.405556:0.033562)0.266667:0.035456,((PA13:0.000001,Bos_taurus:0.000001)1.000000:0.120436,(Lama_glama:0.092094,PA11:0.000001)0.883333:0.038140)0.722222:0.027375)0.238889:0.032920,(PA12:0.019329,Sus_scrofa:0.143478)0.805556:0.024075)0.388889:0.048521)0.483333:0.035104)0.277778:0.029646)0.127778:0.023788)0.691667:0.043473,(Carodnia_vieirai:0.098869,((Loxodonta_africana:0.183155,(Moeritherium_lyonsi:0.134223,Trichechus_manatus:0.182561)0.352778:0.034057)0.558333:0.042819,(PA41:0.000001,(Procavia_capensis:0.000001,PA42:0.000001)1.000000:0.100077)0.880556:0.112960)0.347222:0.043940)0.380556:0.051420)0.130556:0.026036)0.255556:0.021612,(((Protolipterna_ellipsodontoides:0.057145,Didolodus_multicuspis:0.071634)0.605556:0.029722,(Phenacodus_intermedius:0.101442,(Hyopsodus_paulus:0.078664,Protungulatum_donnae:0.153168)0.197222:0.027574)0.111111:0.019631)0.127778:0.018903,Apheliscus_insidiosus:0.108157)0.591667:0.044421)0.161111:0.044039)0.175000:0.036142)0.038889:0.016101,PA5:0.003154)0.025000:0.014642,(((PA8:0.053977,(((PA22:0.000001,(PA21:0.000001,Felis_silvestris:0.000001)0.158333:0.000001)1.000000:0.077276,Canis_lupus:0.064997)1.000000:0.053674,(Sinopa_rapax:0.107614,(Vulpavus_ovatus:0.015745,Vulpavus_profectus:0.028084)1.000000:0.053776)0.430556:0.038047)0.994444:0.056030)0.372222:0.039864,(PA7:0.022208,(Pteropus_giganteus:0.159216,((Icaronycteris_index:0.028310,Onychonycteris_finneyi:0.030166)1.000000:0.076224,((Pteronotus_parnellii:0.075218,(PA17:0.000001,Myotis_lucifugus:0.042438)0.955556:0.018623)0.769444:0.027084,((PA16:0.002378,(Rhinopoma_hardwickii:0.000001,PA20:0.000001)1.000000:0.055181)0.505556:0.012222,((Saccopteryx_bilineata:0.000001,PA19:0.000001)1.000000:0.044106,(PA18:0.014455,Nycteris_thebaica:0.092720)0.727778:0.011518)0.544444:0.021515)0.688889:0.028040)0.991667:0.054199)0.969444:0.048672)1.000000:0.118041)0.286111:0.025114)0.197222:0.039108,(((Galeopterus_variegatus:0.018167,(PA34:0.000001,Cynocephalus_volans:0.000001)1.000000:0.017482)1.000000:0.168912,((PA33:0.015227,(((Tarsius_syrichta:0.136283,(Homo_sapiens:0.162470,(PA36:0.004209,(Saimiri_sciureus:0.000001,PA37:0.000001)1.000000:0.036621)0.986111:0.025818)1.000000:0.050342)0.519444:0.010246,PA35:0.013557)0.838889:0.030721,(Notharctus_tenebrosus:0.138767,Lemur_catta:0.056846)0.480556:0.026056)0.458333:0.013973)1.000000:0.071505,(PA26:0.000001,(Ptilocercus_lowii:0.070438,((PA32:0.000001,Tupaia_glis:0.000001)1.000000:0.031823,PA27:0.014813)1.000000:0.036174)1.000000:0.055333)0.850000:0.028012)0.561111:0.031590)0.997222:0.059544,PA6:0.004873)0.594444:0.025449)0.163889:0.035367)0.088889:0.029598,((((((Amblysomus_hottentotus:0.238638,(Echinops_telfairi:0.000001,PA45:0.000001)1.000000:0.121631)0.841667:0.018702,PA44:0.003020)0.919444:0.010903,PA43:0.003439)0.919444:0.036027,((PA40:0.000001,PA38:0.002438)0.686111:0.050728,Rhynchocyon_cirnei:0.087909)0.205556:0.052437)0.436111:0.045627,Leptictis_dakotensis:0.183847)0.175000:0.039797,(Solenodon_paradoxus:0.165792,(Erinaceus_europaeus:0.124016,(PA23:0.002481,((Sorex_araneus:0.000001,PA25:0.000001)1.000000:0.117073,(PA24:0.008306,Talpa_europaea:0.143045)0.269444:0.022831)0.955556:0.063327)0.588889:0.016461)0.869444:0.042169)0.936111:0.053045)0.105556:0.037271)0.797222:0.027291,PA3:0.002242)0.463889:0.007042,PA1:0.013197)0.891667:0.047653)0.816667:0.046174,((Maelestes_gobiensis:0.098418,(Ukhaatherium_nessovi:0.099510,Zalambdalestes_lechei:0.130143)0.975000:0.058484)0.602778:0.035840,Eomaia_scansoria:0.054300)0.433333:0.040971)1.000000:0.086666,(Zhangheotherium_quinquecuspidens:0.057809,Henkelotherium_guimarotae:0.068008)0.805556:0.038658)0.997222:0.061398)1.000000:0.065303,Haldanodon_exspectatus:0.099776)1.000000:0.078565,Morganucodon_watsoni:0.008634,Morganucodon_oehleri:0.007623);

["dental only" PAs corrected for the O'Leary et al. character ontology: nodal values = "transfer bootstrap expectation" values]
((((Tachyglossus_aculeatus:0.089091,(PA2:0.000001,Ornithorhynchus_anatinus:0.000001)1.000000:0.079871)1.000000:0.136108,(((((Pucadelphys_andinus:0.115891,Didelphis_virginiana:0.052945)0.980556:0.057909,(Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.076726)1.000000:0.066456,((((((((((PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.184910,(Hapalops_elongatus:0.160648,Tamandua_tetradactyla:0.144268)1.000000:0.078499)0.988889:0.057124,(Metacheiromys_marshi:0.117476,Manis_pentadactyla:0.192029)0.961111:0.061191)0.970000:0.044126,Orycteropus_afer:0.232959)0.966204:0.058461,((((Rattus_norvegicus:0.146435,((PA31:0.000001,Ictidomys_tridecemlineatus:0.000001)1.000000:0.060622,((Cavia_porcellus:0.070340,PA28:0.005067)0.616667:0.025921,(PA29:0.003594,(PA30:0.000001,Castor_canadensis:0.000001)1.000000:0.076848)0.951389:0.018578)0.942361:0.034645)0.977315:0.051210)0.957937:0.028559,(Cocomys_lingchaensis:0.073284,Paramys_delicatus:0.052833)0.544444:0.030694)0.950926:0.031304,((Tribosphenomys_minutus:0.119274,Rhombomylus_turpanensis:0.066816)0.236111:0.022397,((Dawsonolagus_antiquus:0.025436,Oryctolagus_cuniculus:0.100239)0.975000:0.033817,Gomphos_elkema:0.080236)1.000000:0.069179)0.956944:0.046631)1.000000:0.090575,((Thomashuxleya_externa_comb:0.156600,((Mesonyx_obtusidens:0.173312,(((((Caperea_marginata:0.171521,(Tursiops_truncatus:0.000001,PA15:0.000001)0.980556:0.169959)0.990278:0.113045,Basilosaurus_cetoides:0.147998)0.864815:0.053288,Artiocetus_clavis:0.124237)0.974306:0.062851,Rodhocetus_balochistanensis:0.103411)0.880556:0.044750,(Archaeotherium_mortoni:0.140562,((PA14:0.000001,Hippopotamus_amphibius:0.000001)1.000000:0.177719,(((PA10:0.013937,((Mesohippus_bairdi:0.191032,Equus_caballus:0.105936)0.572222:0.033170,PA9:0.029627)0.526389:0.033562)0.463889:0.035456,((PA13:0.000001,Bos_taurus:0.000001)1.000000:0.120436,(Lama_glama:0.092094,PA11:0.000001)0.883333:0.038140)0.830556:0.027375)0.642063:0.032920,(PA12:0.019329,Sus_scrofa:0.143478)0.805556:0.024075)0.809877:0.048521)0.874242:0.035104)0.856250:0.029646)0.920833:0.023788)0.965643:0.043473,(Carodnia_vieirai:0.098869,((Loxodonta_africana:0.183155,(Moeritherium_lyonsi:0.134223,Trichechus_manatus:0.182561)0.352778:0.034057)0.740278:0.042819,(PA41:0.000001,(Procavia_capensis:0.000001,PA42:0.000001)1.000000:0.100077)0.940278:0.112960)0.719444:0.043940)0.776389:0.051420)0.847650:0.026036)0.854424:0.021612,(((Protolipterna_ellipsodontoides:0.057145,Didolodus_multicuspis:0.071634)0.605556:0.029722,(Phenacodus_intermedius:0.101442,(Hyopsodus_paulus:0.078664,Protungulatum_donnae:0.153168)0.197222:0.027574)0.415278:0.019631)0.710417:0.018903,Apheliscus_insidiosus:0.108157)0.897222:0.044421)0.810522:0.044039)0.817014:0.036142)0.761970:0.016101,PA5:0.003154)0.754960:0.014642,(((PA8:0.053977,(((PA22:0.000001,(PA21:0.000001,Felis_silvestris:0.000001)0.158333:0.000001)1.000000:0.077276,Canis_lupus:0.064997)1.000000:0.053674,(Sinopa_rapax:0.107614,(Vulpavus_ovatus:0.015745,Vulpavus_profectus:0.028084)1.000000:0.053776)0.715278:0.038047)0.999074:0.056030)0.910317:0.039864,(PA7:0.022208,(Pteropus_giganteus:0.159216,((Icaronycteris_index:0.028310,Onychonycteris_finneyi:0.030166)1.000000:0.076224,((Pteronotus_parnellii:0.075218,(PA17:0.000001,Myotis_lucifugus:0.042438)0.955556:0.018623)0.884722:0.027084,((PA16:0.002378,(Rhinopoma_hardwickii:0.000001,PA20:0.000001)1.000000:0.055181)0.752778:0.012222,((Saccopteryx_bilineata:0.000001,PA19:0.000001)1.000000:0.044106,(PA18:0.014455,Nycteris_thebaica:0.092720)0.727778:0.011518)0.709259:0.021515)0.916204:0.028040)0.998148:0.054199)0.997222:0.048672)1.000000:0.118041)0.945085:0.025114)0.713889:0.039108,(((Galeopterus_variegatus:0.018167,(PA34:0.000001,Cynocephalus_volans:0.000001)1.000000:0.017482)1.000000:0.168912,((PA33:0.015227,(((Tarsius_syrichta:0.136283,(Homo_sapiens:0.162470,(PA36:0.004209,(Saimiri_sciureus:0.000001,PA37:0.000001)1.000000:0.036621)0.993056:0.025818)1.000000:0.050342)0.879861:0.010246,PA35:0.013557)0.942778:0.030721,(Notharctus_tenebrosus:0.138767,Lemur_catta:0.056846)0.480556:0.026056)0.922619:0.013973)1.000000:0.071505,(PA26:0.000001,(Ptilocercus_lowii:0.070438,((PA32:0.000001,Tupaia_glis:0.000001)1.000000:0.031823,PA27:0.014813)1.000000:0.036174)1.000000:0.055333)0.962500:0.028012)0.898718:0.031590)0.999479:0.059544,PA6:0.004873)0.975654:0.025449)0.597721:0.035367)0.598653:0.029598,((((((Amblysomus_hottentotus:0.238638,(Echinops_telfairi:0.000001,PA45:0.000001)1.000000:0.121631)0.920833:0.018702,PA44:0.003020)0.971296:0.010903,PA43:0.003439)0.970833:0.036027,((PA40:0.000001,PA38:0.002438)0.686111:0.050728,Rhynchocyon_cirnei:0.087909)0.445833:0.052437)0.801190:0.045627,Leptictis_dakotensis:0.183847)0.731944:0.039797,(Solenodon_paradoxus:0.165792,(Erinaceus_europaeus:0.124016,(PA23:0.002481,((Sorex_araneus:0.000001,PA25:0.000001)1.000000:0.117073,(PA24:0.008306,Talpa_europaea:0.143045)0.269444:0.022831)0.985185:0.063327)0.893750:0.016461)0.972222:0.042169)0.988426:0.053045)0.517407:0.037271)0.972712:0.027291,PA3:0.002242)0.955382:0.007042,PA1:0.013197)0.984815:0.047653)0.950758:0.046174,((Maelestes_gobiensis:0.098418,(Ukhaatherium_nessovi:0.099510,Zalambdalestes_lechei:0.130143)0.975000:0.058484)0.800000:0.035840,Eomaia_scansoria:0.054300)0.769444:0.040971)1.000000:0.086666,(Zhangheotherium_quinquecuspidens:0.057809,Henkelotherium_guimarotae:0.068008)0.805556:0.038658)0.998889:0.061398)1.000000:0.065303,Haldanodon_exspectatus:0.099776)1.000000:0.078565,Morganucodon_watsoni:0.008634,Morganucodon_oehleri:0.007623);

["dental only" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: nodal values = standard bootstrap values]
(Morganucodon_watsoni:0.008752,(Haldanodon_exspectatus:0.100123,((Ornithorhynchus_anatinus:0.080743,Tachyglossus_aculeatus:0.088416)1.000000:0.135782,((((((Didelphis_virginiana:0.053577,Pucadelphys_andinus:0.114906)1.000000:0.059188,(Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.074525)1.000000:0.063816,(PA3:0.004441,(PA1:0.011203,((PA6:0.035103,(((Pteropus_giganteus:0.168609,((Onychonycteris_finneyi:0.030906,Icaronycteris_index:0.027221)1.000000:0.079678,((Myotis_lucifugus:0.061635,Pteronotus_parnellii:0.076574)0.780556:0.027022,(Nycteris_thebaica:0.093177,(Rhinopoma_hardwickii:0.051089,Saccopteryx_bilineata:0.052974)0.733333:0.024375)0.997222:0.031062)0.969444:0.040036)0.958333:0.045503)1.000000:0.088458,PA8:0.062107)0.013889:0.053675,((Solenodon_paradoxus:0.177668,(Erinaceus_europaeus:0.155046,(Sorex_araneus:0.131361,Talpa_europaea:0.151625)1.000000:0.075335)0.752778:0.036509)0.819444:0.017543,PA7:0.000001)0.513889:0.029817)0.000000:0.031414)0.000000:0.024076,(((PA26:0.003436,(Ptilocercus_lowii:0.070914,(PA27:0.013053,Tupaia_glis:0.031532)0.994444:0.036292)1.000000:0.053988)0.122222:0.027820,((Cynocephalus_volans:0.017457,Galeopterus_variegatus:0.018272)1.000000:0.167320,((Notharctus_tenebrosus:0.135955,Lemur_catta:0.059145)0.330556:0.034917,(PA33:0.000001,(Tarsius_syrichta:0.153260,(Homo_sapiens:0.162609,Saimiri_sciureus:0.066544)1.000000:0.047915)0.316667:0.022254)0.186111:0.005981)1.000000:0.067713)0.994444:0.031978)1.000000:0.092426,((((((Paramys_delicatus:0.053849,Cocomys_lingchaensis:0.072641)0.755556:0.021665,(Rattus_norvegicus:0.147788,(Ictidomys_tridecemlineatus:0.058687,(Castor_canadensis:0.088765,(PA28:0.000001,Cavia_porcellus:0.075771)0.058333:0.018973)0.863889:0.044735)0.813889:0.050095)0.911111:0.030136)0.669444:0.029242,((Rhombomylus_turpanensis:0.081924,(Gomphos_elkema:0.077953,(Oryctolagus_cuniculus:0.099752,Dawsonolagus_antiquus:0.025312)0.975000:0.034770)1.000000:0.066916)0.647222:0.023000,Tribosphenomys_minutus:0.109122)0.769444:0.033317)1.000000:0.098300,(((Leptictis_dakotensis:0.180891,(Sinopa_rapax:0.087192,((Canis_lupus:0.056522,(Felis_silvestris:0.000001,PA21:0.000001)1.000000:0.085133)0.972222:0.066620,(Vulpavus_profectus:0.027163,Vulpavus_ovatus:0.017369)1.000000:0.059322)0.038889:0.027927)0.977778:0.068645)0.000000:0.019060,Protungulatum_donnae:0.137264)0.000000:0.024805,((((Didolodus_multicuspis:0.075196,Protolipterna_ellipsodontoides:0.053755)0.583333:0.034835,(Hyopsodus_paulus:0.103784,Phenacodus_intermedius:0.089316)0.463889:0.025079)0.105556:0.014849,Apheliscus_insidiosus:0.113363)0.230556:0.026443,(((Mesonyx_obtusidens:0.166138,(((Sus_scrofa:0.164880,((PA9:0.025939,(Mesohippus_bairdi:0.188948,Equus_caballus:0.109764)0.727778:0.046280)0.108333:0.027615,((PA10:0.000001,Lama_glama:0.115835)0.077778:0.010271,Bos_taurus:0.130433)0.941667:0.034565)0.097222:0.047078)0.113889:0.040600,Hippopotamus_amphibius:0.188895)0.172222:0.030132,Archaeotherium_mortoni:0.139946)0.058333:0.029009)0.016667:0.019836,(Rodhocetus_balochistanensis:0.114230,(Artiocetus_clavis:0.123765,((Tursiops_truncatus:0.168820,Caperea_marginata:0.168561)1.000000:0.111811,Basilosaurus_cetoides:0.152674)0.536111:0.055586)0.944444:0.055114)0.691667:0.040171)0.888889:0.039260,(Thomashuxleya_externa_comb:0.148611,(Carodnia_vieirai:0.080952,((Trichechus_manatus:0.194255,Loxodonta_africana:0.174999)0.002778:0.036400,Moeritherium_lyonsi:0.138434)0.008333:0.043877)0.005556:0.051639)0.000000:0.037519)0.000000:0.047817)0.000000:0.035153)0.000000:0.030721)0.000000:0.033830,((((PA40:0.000001,(((PA42:0.000001,Procavia_capensis:0.000001)1.000000:0.061668,PA41:0.036882)0.744444:0.144645,PA38:0.000001)0.000000:0.002403)0.000000:0.033362,Rhynchocyon_cirnei:0.128983)0.000000:0.038934,(((Amblysomus_hottentotus:0.250110,PA44:0.000001)0.838889:0.006525,Echinops_telfairi:0.120295)0.536111:0.012497,PA43:0.000001)0.444444:0.055640)0.000000:0.037144,(Orycteropus_afer:0.233441,(((Dasypus_novemcinctus:0.000001,PA39:0.000001)1.000000:0.185160,(Tamandua_tetradactyla:0.142722,Hapalops_elongatus:0.159456)1.000000:0.076155)0.888889:0.054026,(Metacheiromys_marshi:0.116105,Manis_pentadactyla:0.191550)0.991667:0.062652)0.363889:0.035716)0.013889:0.053072)0.000000:0.037393)0.000000:0.028040,PA5:0.001266)0.000000:0.010687)0.000000:0.024726)0.666667:0.030589)0.000000:0.006391)0.000000:0.045706)0.000000:0.049457,(Maelestes_gobiensis:0.099662,(Ukhaatherium_nessovi:0.102444,Zalambdalestes_lechei:0.125821)0.830556:0.057243)0.488889:0.044798)0.005556:0.036996,Eomaia_scansoria:0.051343)0.438889:0.078893,(Henkelotherium_guimarotae:0.068258,Zhangheotherium_quinquecuspidens:0.058109)0.950000:0.044799)0.002778:0.057778)1.000000:0.062189)1.000000:0.076825,Morganucodon_oehleri:0.007453);

["dental only" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: nodal values = "transfer bootstrap expectation" values]
(Morganucodon_watsoni:0.008752,(Haldanodon_exspectatus:0.100123,((Ornithorhynchus_anatinus:0.080743,Tachyglossus_aculeatus:0.088416)1.000000:0.135782,((((((Didelphis_virginiana:0.053577,Pucadelphys_andinus:0.114906)1.000000:0.059188,(Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.074525)1.000000:0.063816,(PA3:0.004441,(PA1:0.011203,((PA6:0.035103,(((Pteropus_giganteus:0.168609,((Onychonycteris_finneyi:0.030906,Icaronycteris_index:0.027221)1.000000:0.079678,((Myotis_lucifugus:0.061635,Pteronotus_parnellii:0.076574)0.780556:0.027022,(Nycteris_thebaica:0.093177,(Rhinopoma_hardwickii:0.051089,Saccopteryx_bilineata:0.052974)0.733333:0.024375)0.998611:0.031062)0.984722:0.040036)0.993056:0.045503)1.000000:0.088458,PA8:0.062107)0.876736:0.053675,((Solenodon_paradoxus:0.177668,(Erinaceus_europaeus:0.155046,(Sorex_araneus:0.131361,Talpa_europaea:0.151625)1.000000:0.075335)0.876389:0.036509)0.933333:0.017543,PA7:0.000001)0.843056:0.029817)0.583547:0.031414)0.541865:0.024076,(((PA26:0.003436,(Ptilocercus_lowii:0.070914,(PA27:0.013053,Tupaia_glis:0.031532)0.994444:0.036292)1.000000:0.053988)0.707407:0.027820,((Cynocephalus_volans:0.017457,Galeopterus_variegatus:0.018272)1.000000:0.167320,((Notharctus_tenebrosus:0.135955,Lemur_catta:0.059145)0.330556:0.034917,(PA33:0.000001,(Tarsius_syrichta:0.153260,(Homo_sapiens:0.162609,Saimiri_sciureus:0.066544)1.000000:0.047915)0.658333:0.022254)0.510185:0.005981)1.000000:0.067713)0.998413:0.031978)1.000000:0.092426,((((((Paramys_delicatus:0.053849,Cocomys_lingchaensis:0.072641)0.755556:0.021665,(Rattus_norvegicus:0.147788,(Ictidomys_tridecemlineatus:0.058687,(Castor_canadensis:0.088765,(PA28:0.000001,Cavia_porcellus:0.075771)0.058333:0.018973)0.930556:0.044735)0.937037:0.050095)0.972917:0.030136)0.894907:0.029242,((Rhombomylus_turpanensis:0.081924,(Gomphos_elkema:0.077953,(Oryctolagus_cuniculus:0.099752,Dawsonolagus_antiquus:0.025312)0.975000:0.034770)1.000000:0.066916)0.882407:0.023000,Tribosphenomys_minutus:0.109122)0.940278:0.033317)1.000000:0.098300,(((Leptictis_dakotensis:0.180891,(Sinopa_rapax:0.087192,((Canis_lupus:0.056522,(Felis_silvestris:0.000001,PA21:0.000001)1.000000:0.085133)0.986111:0.066620,(Vulpavus_profectus:0.027163,Vulpavus_ovatus:0.017369)1.000000:0.059322)0.754167:0.027927)0.986667:0.068645)0.822222:0.019060,Protungulatum_donnae:0.137264)0.705952:0.024805,((((Didolodus_multicuspis:0.075196,Protolipterna_ellipsodontoides:0.053755)0.583333:0.034835,(Hyopsodus_paulus:0.103784,Phenacodus_intermedius:0.089316)0.463889:0.025079)0.513889:0.014849,Apheliscus_insidiosus:0.113363)0.738194:0.026443,(((Mesonyx_obtusidens:0.166138,(((Sus_scrofa:0.164880,((PA9:0.025939,(Mesohippus_bairdi:0.188948,Equus_caballus:0.109764)0.727778:0.046280)0.484722:0.027615,((PA10:0.000001,Lama_glama:0.115835)0.077778:0.010271,Bos_taurus:0.130433)0.968056:0.034565)0.536667:0.047078)0.534722:0.040600,Hippopotamus_amphibius:0.188895)0.522222:0.030132,Archaeotherium_mortoni:0.139946)0.481597:0.029009)0.555864:0.019836,(Rodhocetus_balochistanensis:0.114230,(Artiocetus_clavis:0.123765,((Tursiops_truncatus:0.168820,Caperea_marginata:0.168561)1.000000:0.111811,Basilosaurus_cetoides:0.152674)0.768056:0.055586)0.981481:0.055114)0.921528:0.040171)0.963294:0.039260,(Thomashuxleya_externa_comb:0.148611,(Carodnia_vieirai:0.080952,((Trichechus_manatus:0.194255,Loxodonta_africana:0.174999)0.002778:0.036400,Moeritherium_lyonsi:0.138434)0.237500:0.043877)0.328704:0.051639)0.373611:0.037519)0.720322:0.047817)0.649074:0.035153)0.530903:0.030721)0.512184:0.033830,((((PA40:0.000001,(((PA42:0.000001,Procavia_capensis:0.000001)1.000000:0.061668,PA41:0.036882)0.872222:0.144645,PA38:0.000001)0.581481:0.002403)0.436806:0.033362,Rhynchocyon_cirnei:0.128983)0.349444:0.038934,(((Amblysomus_hottentotus:0.250110,PA44:0.000001)0.838889:0.006525,Echinops_telfairi:0.120295)0.766667:0.012497,PA43:0.000001)0.814815:0.055640)0.478704:0.037144,(Orycteropus_afer:0.233441,(((Dasypus_novemcinctus:0.000001,PA39:0.000001)1.000000:0.185160,(Tamandua_tetradactyla:0.142722,Hapalops_elongatus:0.159456)1.000000:0.076155)0.925926:0.054026,(Metacheiromys_marshi:0.116105,Manis_pentadactyla:0.191550)0.991667:0.062652)0.746667:0.035716)0.626852:0.053072)0.749132:0.037393)0.475000:0.028040,PA5:0.001266)0.464987:0.010687)0.477419:0.024726)0.922049:0.030589)0.851296:0.006391)0.841270:0.045706)0.703056:0.049457,(Maelestes_gobiensis:0.099662,(Ukhaatherium_nessovi:0.102444,Zalambdalestes_lechei:0.125821)0.830556:0.057243)0.743056:0.044798)0.748016:0.036996,Eomaia_scansoria:0.051343)0.813889:0.078893,(Henkelotherium_guimarotae:0.068258,Zhangheotherium_quinquecuspidens:0.058109)0.950000:0.044799)0.501389:0.057778)1.000000:0.062189)1.000000:0.076825,Morganucodon_oehleri:0.007453);

["dental only" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: nodal values = standard bootstrap values]
((Haldanodon_exspectatus:0.100434,((Tachyglossus_aculeatus:0.088937,Ornithorhynchus_anatinus:0.080010)1.000000:0.136503,((Henkelotherium_guimarotae:0.069217,Zhangheotherium_quinquecuspidens:0.058144)0.829365:0.039308,((((Didelphis_virginiana:0.053619,Pucadelphys_andinus:0.115766)0.996032:0.060769,(Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.074092)1.000000:0.068261,(PA1:0.011011,(PA3:0.005071,(((((Talpa_europaea:0.135126,Sorex_araneus:0.143561)0.559524:0.048975,Amblysomus_hottentotus:0.239776)0.900794:0.061935,(Erinaceus_europaeus:0.124896,Echinops_telfairi:0.151453)0.904762:0.048121)0.543651:0.027448,(Solenodon_paradoxus:0.167885,PA7:0.035650)0.250000:0.028806)0.373016:0.048271,((PA5:0.000879,((Orycteropus_afer:0.244172,(((Tamandua_tetradactyla:0.145071,Hapalops_elongatus:0.158854)1.000000:0.078170,(PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.187538)0.956349:0.054950,(Manis_pentadactyla:0.192582,Metacheiromys_marshi:0.117944)0.964286:0.061199)0.912698:0.038890)0.888889:0.057391,((((((Sinopa_rapax:0.086945,((Vulpavus_profectus:0.027190,Vulpavus_ovatus:0.017205)1.000000:0.059725,(Canis_lupus:0.066871,Felis_silvestris:0.098071)1.000000:0.055077)0.888889:0.027665)1.000000:0.065042,Protungulatum_donnae:0.130585)0.222222:0.018854,Leptictis_dakotensis:0.186925)0.142857:0.033105,((Thomashuxleya_externa_comb:0.165505,((((Archaeotherium_mortoni:0.142199,((((Mesohippus_bairdi:0.164285,Equus_caballus:0.132200)0.976190:0.066378,(Bos_taurus:0.125307,Lama_glama:0.123049)1.000000:0.051489)0.357143:0.035583,Sus_scrofa:0.173838)0.373016:0.034812,Hippopotamus_amphibius:0.195108)0.341270:0.027243)0.396825:0.029015,Mesonyx_obtusidens:0.167620)0.202381:0.018335,((Artiocetus_clavis:0.121991,(Basilosaurus_cetoides:0.154805,(Caperea_marginata:0.169745,Tursiops_truncatus:0.171195)1.000000:0.112649)0.646825:0.054980)0.948413:0.059499,Rodhocetus_balochistanensis:0.114209)0.476190:0.041021)0.825397:0.033668,((Moeritherium_lyonsi:0.141135,(Loxodonta_africana:0.176626,Trichechus_manatus:0.195070)0.500000:0.035935)0.698413:0.041227,Carodnia_vieirai:0.088362)0.904762:0.068397)0.309524:0.028932)0.630952:0.035898,(((Didolodus_multicuspis:0.073909,Protolipterna_ellipsodontoides:0.054382)0.642857:0.034501,(Hyopsodus_paulus:0.103920,Phenacodus_intermedius:0.089264)0.551587:0.025782)0.349206:0.013012,Apheliscus_insidiosus:0.117037)0.579365:0.026756)0.246032:0.035884)0.123016:0.029210,((Tribosphenomys_minutus:0.109275,(((Oryctolagus_cuniculus:0.099793,Dawsonolagus_antiquus:0.025405)0.972222:0.035488,Gomphos_elkema:0.077039)1.000000:0.070197,Rhombomylus_turpanensis:0.079305)0.599206:0.023127)0.869048:0.034496,((Rattus_norvegicus:0.152461,(Ictidomys_tridecemlineatus:0.065672,(Castor_canadensis:0.086918,Cavia_porcellus:0.090535)0.940476:0.048987)0.571429:0.045969)0.361111:0.027322,(Cocomys_lingchaensis:0.075691,Paramys_delicatus:0.052172)0.404762:0.020557)0.710317:0.031285)1.000000:0.093493)0.059524:0.026126,(Procavia_capensis:0.206409,((PA38:0.000826,PA40:0.001625)0.948413:0.018393,Rhynchocyon_cirnei:0.110025)0.865079:0.042043)0.734127:0.075748)0.261905:0.027208)0.242063:0.015170)0.226190:0.019497,(((Cynocephalus_volans:0.016723,Galeopterus_variegatus:0.019115)1.000000:0.171279,((PA26:0.003670,((Notharctus_tenebrosus:0.127942,Lemur_catta:0.067465)0.690476:0.034218,((Saimiri_sciureus:0.071827,Homo_sapiens:0.158993)1.000000:0.054014,Tarsius_syrichta:0.142212)0.726190:0.027257)1.000000:0.062549)0.384921:0.020532,(Tupaia_glis:0.066112,Ptilocercus_lowii:0.073342)1.000000:0.064035)0.531746:0.032476)0.992063:0.059435,((Pteropus_giganteus:0.164208,((Icaronycteris_index:0.026733,Onychonycteris_finneyi:0.031489)1.000000:0.069867,((Myotis_lucifugus:0.062131,Pteronotus_parnellii:0.076600)0.785714:0.027173,((Rhinopoma_hardwickii:0.052508,Saccopteryx_bilineata:0.052152)0.793651:0.025768,Nycteris_thebaica:0.092621)1.000000:0.031751)1.000000:0.046569)0.984127:0.051937)1.000000:0.126883,PA6:0.015514)0.063492:0.019194)0.158730:0.041092)0.051587:0.025108)0.603175:0.024828)0.357143:0.006553)0.873016:0.046644)0.734127:0.040744,((Eomaia_scansoria:0.051703,Maelestes_gobiensis:0.107880)0.281746:0.025615,(Zalambdalestes_lechei:0.131544,Ukhaatherium_nessovi:0.103368)0.964286:0.060157)0.404762:0.044590)1.000000:0.093006)1.000000:0.061093)1.000000:0.066899)1.000000:0.079158,Morganucodon_watsoni:0.008331,Morganucodon_oehleri:0.008015);

["dental only" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: nodal values = "transfer bootstrap expectation" values]
((Haldanodon_exspectatus:0.100434,((Tachyglossus_aculeatus:0.088937,Ornithorhynchus_anatinus:0.080010)1.000000:0.136503,((Henkelotherium_guimarotae:0.069217,Zhangheotherium_quinquecuspidens:0.058144)0.829365:0.039308,((((Didelphis_virginiana:0.053619,Pucadelphys_andinus:0.115766)0.996032:0.060769,(Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.074092)1.000000:0.068261,(PA1:0.011011,(PA3:0.005071,(((((Talpa_europaea:0.135126,Sorex_araneus:0.143561)0.559524:0.048975,Amblysomus_hottentotus:0.239776)0.950397:0.061935,(Erinaceus_europaeus:0.124896,Echinops_telfairi:0.151453)0.904762:0.048121)0.812500:0.027448,(Solenodon_paradoxus:0.167885,PA7:0.035650)0.250000:0.028806)0.830688:0.048271,((PA5:0.000879,((Orycteropus_afer:0.244172,(((Tamandua_tetradactyla:0.145071,Hapalops_elongatus:0.158854)1.000000:0.078170,(PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.187538)0.970899:0.054950,(Manis_pentadactyla:0.192582,Metacheiromys_marshi:0.117944)0.964286:0.061199)0.982540:0.038890)0.981481:0.057391,((((((Sinopa_rapax:0.086945,((Vulpavus_profectus:0.027190,Vulpavus_ovatus:0.017205)1.000000:0.059725,(Canis_lupus:0.066871,Felis_silvestris:0.098071)1.000000:0.055077)0.962963:0.027665)1.000000:0.065042,Protungulatum_donnae:0.130585)0.844444:0.018854,Leptictis_dakotensis:0.186925)0.759259:0.033105,((Thomashuxleya_externa_comb:0.165505,((((Archaeotherium_mortoni:0.142199,((((Mesohippus_bairdi:0.164285,Equus_caballus:0.132200)0.976190:0.066378,(Bos_taurus:0.125307,Lama_glama:0.123049)1.000000:0.051489)0.626984:0.035583,Sus_scrofa:0.173838)0.694444:0.034812,Hippopotamus_amphibius:0.195108)0.726190:0.027243)0.726852:0.029015,Mesonyx_obtusidens:0.167620)0.693878:0.018335,((Artiocetus_clavis:0.121991,(Basilosaurus_cetoides:0.154805,(Caperea_marginata:0.169745,Tursiops_truncatus:0.171195)1.000000:0.112649)0.823413:0.054980)0.974868:0.059499,Rodhocetus_balochistanensis:0.114209)0.859127:0.041021)0.949405:0.033668,((Moeritherium_lyonsi:0.141135,(Loxodonta_africana:0.176626,Trichechus_manatus:0.195070)0.500000:0.035935)0.839286:0.041227,Carodnia_vieirai:0.088362)0.960317:0.068397)0.928323:0.028932)0.950047:0.035898,(((Didolodus_multicuspis:0.073909,Protolipterna_ellipsodontoides:0.054382)0.642857:0.034501,(Hyopsodus_paulus:0.103920,Phenacodus_intermedius:0.089264)0.551587:0.025782)0.710317:0.013012,Apheliscus_insidiosus:0.117037)0.878968:0.026756)0.867965:0.035884)0.772441:0.029210,((Tribosphenomys_minutus:0.109275,(((Oryctolagus_cuniculus:0.099793,Dawsonolagus_antiquus:0.025405)0.972222:0.035488,Gomphos_elkema:0.077039)1.000000:0.070197,Rhombomylus_turpanensis:0.079305)0.866402:0.023127)0.965278:0.034496,((Rattus_norvegicus:0.152461,(Ictidomys_tridecemlineatus:0.065672,(Castor_canadensis:0.086918,Cavia_porcellus:0.090535)0.940476:0.048987)0.785714:0.045969)0.711640:0.027322,(Cocomys_lingchaensis:0.075691,Paramys_delicatus:0.052172)0.404762:0.020557)0.886508:0.031285)1.000000:0.093493)0.833929:0.026126,(Procavia_capensis:0.206409,((PA38:0.000826,PA40:0.001625)0.948413:0.018393,Rhynchocyon_cirnei:0.110025)0.920635:0.042043)0.896825:0.075748)0.877435:0.027208)0.856958:0.015170)0.851285:0.019497,(((Cynocephalus_volans:0.016723,Galeopterus_variegatus:0.019115)1.000000:0.171279,((PA26:0.003670,((Notharctus_tenebrosus:0.127942,Lemur_catta:0.067465)0.690476:0.034218,((Saimiri_sciureus:0.071827,Homo_sapiens:0.158993)1.000000:0.054014,Tarsius_syrichta:0.142212)0.863095:0.027257)1.000000:0.062549)0.876984:0.020532,(Tupaia_glis:0.066112,Ptilocercus_lowii:0.073342)1.000000:0.064035)0.866213:0.032476)0.998236:0.059435,((Pteropus_giganteus:0.164208,((Icaronycteris_index:0.026733,Onychonycteris_finneyi:0.031489)1.000000:0.069867,((Myotis_lucifugus:0.062131,Pteronotus_parnellii:0.076600)0.785714:0.027173,((Rhinopoma_hardwickii:0.052508,Saccopteryx_bilineata:0.052152)0.793651:0.025768,Nycteris_thebaica:0.092621)1.000000:0.031751)1.000000:0.046569)0.997354:0.051937)1.000000:0.126883,PA6:0.015514)0.882937:0.019194)0.653880:0.041092)0.736715:0.025108)0.942956:0.024828)0.942328:0.006553)0.980159:0.046644)0.922619:0.040744,((Eomaia_scansoria:0.051703,Maelestes_gobiensis:0.107880)0.281746:0.025615,(Zalambdalestes_lechei:0.131544,Ukhaatherium_nessovi:0.103368)0.964286:0.060157)0.755291:0.044590)1.000000:0.093006)1.000000:0.061093)1.000000:0.066899)1.000000:0.079158,Morganucodon_watsoni:0.008331,Morganucodon_oehleri:0.008015);

["max preservation" PAs corrected for the O'Leary et al. character ontology: nodal values = standard bootstrap values]
((((Henkelotherium_guimarotae:0.079206,Zhangheotherium_quinquecuspidens:0.054705)0.920000:0.060398,(((PA2:0.000001,Ornithorhynchus_anatinus:0.000001)1.000000:0.075562,Tachyglossus_aculeatus:0.078816)1.000000:0.076046,(PA1:0.000001,((PA3:0.000001,(((PA6:0.000001,((PA26:0.000001,((PA33:0.000001,((PA35:0.009269,((Homo_sapiens:0.151166,(PA36:0.001887,(Saimiri_sciureus:0.000001,PA37:0.000001)1.000000:0.044673)0.696667:0.014753)1.000000:0.054040,Tarsius_syrichta:0.113416)0.623333:0.017963)0.886667:0.037275,(Notharctus_tenebrosus:0.152884,Lemur_catta:0.042862)0.850000:0.022585)0.890000:0.029480)1.000000:0.058996,(Galeopterus_variegatus:0.018942,(Cynocephalus_volans:0.000001,PA34:0.000001)1.000000:0.015687)1.000000:0.143540)0.983333:0.061901)0.873333:0.031244,((PA27:0.007030,(PA32:0.000001,Tupaia_glis:0.000001)1.000000:0.047284)0.996667:0.022429,Ptilocercus_lowii:0.065283)1.000000:0.054917)1.000000:0.095637)0.726667:0.029301,((PA7:0.000204,((Metacheiromys_marshi:0.144166,Manis_pentadactyla:0.170877)0.990000:0.108059,((((Vulpavus_profectus:0.026233,Vulpavus_ovatus:0.016871)1.000000:0.051527,Sinopa_rapax:0.110374)0.963333:0.073068,(Canis_lupus:0.058752,(PA21:0.000001,(PA22:0.000001,Felis_silvestris:0.000001)1.000000:0.028142)1.000000:0.047310)0.976667:0.042376)0.966667:0.061079,((((((Rhinopoma_hardwickii:0.000001,PA20:0.000001)1.000000:0.050228,(((Pteronotus_parnellii:0.072424,(PA17:0.000001,Myotis_lucifugus:0.053865)0.983333:0.008632)0.863333:0.019346,((Nycteris_thebaica:0.090812,PA18:0.006039)0.900000:0.011413,(PA19:0.000001,Saccopteryx_bilineata:0.000001)1.000000:0.045366)1.000000:0.027376)0.906667:0.031588,PA16:0.000001)0.853333:0.015861)0.896667:0.040489,(Icaronycteris_index:0.022431,Onychonycteris_finneyi:0.033276)1.000000:0.119141)0.483333:0.023934,Pteropus_giganteus:0.162039)1.000000:0.117859,(PA9:0.006783,((Mesohippus_bairdi:0.219184,Equus_caballus:0.083793)0.630000:0.047844,(PA10:0.008352,(Lama_glama:0.087411,(PA11:0.000001,((Bos_taurus:0.037883,(((Tursiops_truncatus:0.000001,PA15:0.000001)1.000000:0.124755,Caperea_marginata:0.155941)1.000000:0.237525,PA13:0.000001)0.313333:0.016449)0.406667:0.053333,((PA14:0.000001,Hippopotamus_amphibius:0.000001)1.000000:0.199581,(Sus_scrofa:0.153521,PA12:0.000001)0.440000:0.010258)0.370000:0.034373)0.383333:0.050044)0.356667:0.034646)0.380000:0.039608)0.393333:0.055511)0.233333:0.032616)0.270000:0.091480)0.190000:0.055703,PA8:0.004406)0.183333:0.046886)0.173333:0.027650)0.100000:0.045738)0.073333:0.038566,(Solenodon_paradoxus:0.112446,(PA23:0.000001,((Talpa_europaea:0.122313,(PA25:0.000001,Sorex_araneus:0.000001)1.000000:0.065857)1.000000:0.065345,(Erinaceus_europaeus:0.102375,PA24:0.000001)0.636667:0.051300)0.960000:0.058429)0.900000:0.045225)1.000000:0.082013)0.120000:0.046805)0.106667:0.030991,((((Rhynchocyon_cirnei:0.095238,(PA43:0.011842,((Echinops_telfairi:0.000001,PA45:0.000001)1.000000:0.129234,(PA44:0.000119,Amblysomus_hottentotus:0.238413)0.656667:0.028976)0.720000:0.032578)0.496667:0.026029)0.983333:0.049438,((((PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.160162,(Hapalops_elongatus:0.181651,Tamandua_tetradactyla:0.116039)1.000000:0.082674)0.886667:0.085159,PA38:0.000001)0.583333:0.024977,(((Trichechus_manatus:0.174684,((Moeritherium_lyonsi:0.189667,Loxodonta_africana:0.138029)0.376667:0.070936,((PA42:0.000001,Procavia_capensis:0.000001)1.000000:0.105581,PA41:0.000001)0.756667:0.061071)0.143333:0.021994)0.476667:0.111275,PA40:0.001992)0.430000:0.018556,Orycteropus_afer:0.220239)0.180000:0.013959)0.163333:0.028173)0.303333:0.078387,((((Rhombomylus_turpanensis:0.083287,(Gomphos_elkema:0.079178,(Dawsonolagus_antiquus:0.025282,Oryctolagus_cuniculus:0.095481)0.983333:0.033853)1.000000:0.063241)0.666667:0.021208,Tribosphenomys_minutus:0.112892)0.783333:0.029886,((Paramys_delicatus:0.052155,Cocomys_lingchaensis:0.072878)0.773333:0.028356,(Rattus_norvegicus:0.117649,(PA28:0.009623,(((PA31:0.000001,Ictidomys_tridecemlineatus:0.000001)1.000000:0.075110,(PA29:0.000001,(Castor_canadensis:0.000001,PA30:0.000001)1.000000:0.072314)0.980000:0.022203)0.543333:0.023740,Cavia_porcellus:0.091722)0.240000:0.007136)0.906667:0.058070)0.970000:0.051812)0.650000:0.022826)1.000000:0.098075,((Protungulatum_donnae:0.130515,((Zalambdalestes_lechei:0.126944,((Eomaia_scansoria:0.083741,Maelestes_gobiensis:0.076934)0.506667:0.052336,Ukhaatherium_nessovi:0.106106)0.210000:0.031660)0.696667:0.066997,Leptictis_dakotensis:0.144122)0.293333:0.032283)0.126667:0.036597,((Mesonyx_obtusidens:0.130218,((Rodhocetus_balochistanensis:0.098428,(Basilosaurus_cetoides:0.189783,Artiocetus_clavis:0.130712)0.330000:0.034589)0.256667:0.048814,Archaeotherium_mortoni:0.116255)0.190000:0.042004)0.396667:0.089689,((Carodnia_vieirai:0.117483,Thomashuxleya_externa_comb:0.123977)0.613333:0.067737,(((Didolodus_multicuspis:0.074080,Protolipterna_ellipsodontoides:0.053353)0.596667:0.029170,(Hyopsodus_paulus:0.098870,Phenacodus_intermedius:0.091614)0.456667:0.025479)0.143333:0.012385,Apheliscus_insidiosus:0.116932)0.236667:0.016793)0.180000:0.023628)0.046667:0.040195)0.026667:0.032825)0.016667:0.038454)0.030000:0.035080,PA5:0.005703)0.030000:0.020019)0.310000:0.047299)0.310000:0.034460,((Didelphis_virginiana:0.039477,Pucadelphys_andinus:0.128845)1.000000:0.060268,(Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.060931)1.000000:0.085209)0.463333:0.066818)0.463333:0.070761)0.996667:0.096670)1.000000:0.103603,Haldanodon_exspectatus:0.097178)1.000000:0.077575,Morganucodon_watsoni:0.007354,Morganucodon_oehleri:0.009061);

["max preservation" PAs corrected for the O'Leary et al. character ontology: nodal values = "transfer bootstrap expectation" values]
((((Henkelotherium_guimarotae:0.079206,Zhangheotherium_quinquecuspidens:0.054705)0.920000:0.060398,(((PA2:0.000001,Ornithorhynchus_anatinus:0.000001)1.000000:0.075562,Tachyglossus_aculeatus:0.078816)1.000000:0.076046,(PA1:0.000001,((PA3:0.000001,(((PA6:0.000001,((PA26:0.000001,((PA33:0.000001,((PA35:0.009269,((Homo_sapiens:0.151166,(PA36:0.001887,(Saimiri_sciureus:0.000001,PA37:0.000001)1.000000:0.044673)0.848333:0.014753)1.000000:0.054040,Tarsius_syrichta:0.113416)0.905833:0.017963)0.956667:0.037275,(Notharctus_tenebrosus:0.152884,Lemur_catta:0.042862)0.850000:0.022585)0.984286:0.029480)1.000000:0.058996,(Galeopterus_variegatus:0.018942,(Cynocephalus_volans:0.000001,PA34:0.000001)1.000000:0.015687)1.000000:0.143540)0.995455:0.061901)0.985278:0.031244,((PA27:0.007030,(PA32:0.000001,Tupaia_glis:0.000001)1.000000:0.047284)0.998333:0.022429,Ptilocercus_lowii:0.065283)1.000000:0.054917)1.000000:0.095637)0.983922:0.029301,((PA7:0.000204,((Metacheiromys_marshi:0.144166,Manis_pentadactyla:0.170877)0.990000:0.108059,((((Vulpavus_profectus:0.026233,Vulpavus_ovatus:0.016871)1.000000:0.051527,Sinopa_rapax:0.110374)0.981667:0.073068,(Canis_lupus:0.058752,(PA21:0.000001,(PA22:0.000001,Felis_silvestris:0.000001)1.000000:0.028142)1.000000:0.047310)0.992222:0.042376)0.983333:0.061079,((((((Rhinopoma_hardwickii:0.000001,PA20:0.000001)1.000000:0.050228,(((Pteronotus_parnellii:0.072424,(PA17:0.000001,Myotis_lucifugus:0.053865)0.983333:0.008632)0.925000:0.019346,((Nycteris_thebaica:0.090812,PA18:0.006039)0.900000:0.011413,(PA19:0.000001,Saccopteryx_bilineata:0.000001)1.000000:0.045366)1.000000:0.027376)0.968889:0.031588,PA16:0.000001)0.965714:0.015861)0.977407:0.040489,(Icaronycteris_index:0.022431,Onychonycteris_finneyi:0.033276)1.000000:0.119141)0.953030:0.023934,Pteropus_giganteus:0.162039)1.000000:0.117859,(PA9:0.006783,((Mesohippus_bairdi:0.219184,Equus_caballus:0.083793)0.630000:0.047844,(PA10:0.008352,(Lama_glama:0.087411,(PA11:0.000001,((Bos_taurus:0.037883,(((Tursiops_truncatus:0.000001,PA15:0.000001)1.000000:0.124755,Caperea_marginata:0.155941)1.000000:0.237525,PA13:0.000001)0.771111:0.016449)0.706667:0.053333,((PA14:0.000001,Hippopotamus_amphibius:0.000001)1.000000:0.199581,(Sus_scrofa:0.153521,PA12:0.000001)0.440000:0.010258)0.586667:0.034373)0.734583:0.050044)0.742963:0.034646)0.785333:0.039608)0.811818:0.055511)0.806154:0.032616)0.834762:0.091480)0.780988:0.055703,PA8:0.004406)0.793214:0.046886)0.845429:0.027650)0.835405:0.045738)0.832018:0.038566,(Solenodon_paradoxus:0.112446,(PA23:0.000001,((Talpa_europaea:0.122313,(PA25:0.000001,Sorex_araneus:0.000001)1.000000:0.065857)1.000000:0.065345,(Erinaceus_europaeus:0.102375,PA24:0.000001)0.636667:0.051300)0.981667:0.058429)0.980000:0.045225)1.000000:0.082013)0.878593:0.046805)0.857037:0.030991,((((Rhynchocyon_cirnei:0.095238,(PA43:0.011842,((Echinops_telfairi:0.000001,PA45:0.000001)1.000000:0.129234,(PA44:0.000119,Amblysomus_hottentotus:0.238413)0.656667:0.028976)0.851111:0.032578)0.874167:0.026029)0.996667:0.049438,((((PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.160162,(Hapalops_elongatus:0.181651,Tamandua_tetradactyla:0.116039)1.000000:0.082674)0.925556:0.085159,PA38:0.000001)0.863333:0.024977,(((Trichechus_manatus:0.174684,((Moeritherium_lyonsi:0.189667,Loxodonta_africana:0.138029)0.376667:0.070936,((PA42:0.000001,Procavia_capensis:0.000001)1.000000:0.105581,PA41:0.000001)0.878333:0.061071)0.710000:0.021994)0.892667:0.111275,PA40:0.001992)0.897222:0.018556,Orycteropus_afer:0.220239)0.827619:0.013959)0.760000:0.028173)0.922222:0.078387,((((Rhombomylus_turpanensis:0.083287,(Gomphos_elkema:0.079178,(Dawsonolagus_antiquus:0.025282,Oryctolagus_cuniculus:0.095481)0.983333:0.033853)1.000000:0.063241)0.888889:0.021208,Tribosphenomys_minutus:0.112892)0.945000:0.029886,((Paramys_delicatus:0.052155,Cocomys_lingchaensis:0.072878)0.773333:0.028356,(Rattus_norvegicus:0.117649,(PA28:0.009623,(((PA31:0.000001,Ictidomys_tridecemlineatus:0.000001)1.000000:0.075110,(PA29:0.000001,(Castor_canadensis:0.000001,PA30:0.000001)1.000000:0.072314)0.990000:0.022203)0.776667:0.023740,Cavia_porcellus:0.091722)0.829333:0.007136)0.983889:0.058070)0.995238:0.051812)0.924815:0.022826)1.000000:0.098075,((Protungulatum_donnae:0.130515,((Zalambdalestes_lechei:0.126944,((Eomaia_scansoria:0.083741,Maelestes_gobiensis:0.076934)0.506667:0.052336,Ukhaatherium_nessovi:0.106106)0.456667:0.031660)0.891111:0.066997,Leptictis_dakotensis:0.144122)0.758333:0.032283)0.638667:0.036597,((Mesonyx_obtusidens:0.130218,((Rodhocetus_balochistanensis:0.098428,(Basilosaurus_cetoides:0.189783,Artiocetus_clavis:0.130712)0.330000:0.034589)0.356667:0.048814,Archaeotherium_mortoni:0.116255)0.361111:0.042004)0.573333:0.089689,((Carodnia_vieirai:0.117483,Thomashuxleya_externa_comb:0.123977)0.613333:0.067737,(((Didolodus_multicuspis:0.074080,Protolipterna_ellipsodontoides:0.053353)0.596667:0.029170,(Hyopsodus_paulus:0.098870,Phenacodus_intermedius:0.091614)0.456667:0.025479)0.537778:0.012385,Apheliscus_insidiosus:0.116932)0.687500:0.016793)0.726667:0.023628)0.596970:0.040195)0.583333:0.032825)0.702187:0.038454)0.760196:0.035080,PA5:0.005703)0.760962:0.020019)0.793590:0.047299)0.776944:0.034460,((Didelphis_virginiana:0.039477,Pucadelphys_andinus:0.128845)1.000000:0.060268,(Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.060931)1.000000:0.085209)0.740417:0.066818)0.774286:0.070761)0.998333:0.096670)1.000000:0.103603,Haldanodon_exspectatus:0.097178)1.000000:0.077575,Morganucodon_watsoni:0.007354,Morganucodon_oehleri:0.009061);

["max preservation" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: nodal values = standard bootstrap values]
(((((Eomaia_scansoria:0.050363,(Maelestes_gobiensis:0.094181,(Zalambdalestes_lechei:0.130308,Ukhaatherium_nessovi:0.098018)0.830556:0.061090)0.488889:0.039351)0.669444:0.042127,(((PA1:0.000001,(Ornithorhynchus_anatinus:0.081211,Tachyglossus_aculeatus:0.084164)1.000000:0.137421)0.555556:0.053505,((((PA6:0.000001,((Ptilocercus_lowii:0.065217,(PA27:0.005678,Tupaia_glis:0.048324)0.994444:0.023286)1.000000:0.055449,((((PA33:0.000001,((Notharctus_tenebrosus:0.117803,Tarsius_syrichta:0.129514)0.669444:0.044953,Lemur_catta:0.067176)0.580556:0.025893)0.563889:0.004696,(Saimiri_sciureus:0.066634,Homo_sapiens:0.157486)1.000000:0.055836)1.000000:0.057510,(Galeopterus_variegatus:0.018979,Cynocephalus_volans:0.016461)1.000000:0.143761)0.994444:0.059195,PA26:0.000001)0.877778:0.030960)1.000000:0.095500)0.769444:0.025985,(((((Sorex_araneus:0.131111,Talpa_europaea:0.149889)1.000000:0.072061,Erinaceus_europaeus:0.156707)0.752778:0.037278,Solenodon_paradoxus:0.173395)0.819444:0.045872,PA7:0.000001)0.513889:0.037614,(((Metacheiromys_marshi:0.136425,Manis_pentadactyla:0.180535)0.991667:0.107220,(((Felis_silvestris:0.032719,PA21:0.000001)1.000000:0.045998,Canis_lupus:0.059355)0.972222:0.048479,(Sinopa_rapax:0.109679,(Vulpavus_profectus:0.026275,Vulpavus_ovatus:0.016773)1.000000:0.051105)0.961111:0.065954)0.977778:0.050752)0.133333:0.017507,(PA8:0.009764,((Pteropus_giganteus:0.177525,((Icaronycteris_index:0.025653,Onychonycteris_finneyi:0.031331)1.000000:0.094601,((Pteronotus_parnellii:0.074717,Myotis_lucifugus:0.061565)0.780556:0.026429,(Nycteris_thebaica:0.094885,(Saccopteryx_bilineata:0.051306,Rhinopoma_hardwickii:0.050841)0.733333:0.022187)0.997222:0.029776)0.969444:0.030646)0.958333:0.040086)1.000000:0.104906,(PA9:0.007741,(((Sus_scrofa:0.151556,(((Mesonyx_obtusidens:0.149841,((Basilosaurus_cetoides:0.154393,(Artiocetus_clavis:0.108897,(Caperea_marginata:0.162523,Tursiops_truncatus:0.158446)1.000000:0.100905)0.425000:0.039164)0.944444:0.057662,Rodhocetus_balochistanensis:0.099855)0.691667:0.036812)0.569444:0.045237,Archaeotherium_mortoni:0.118788)0.755556:0.056904,Hippopotamus_amphibius:0.152424)0.619444:0.060410)0.561111:0.041576,(PA10:0.000001,(Bos_taurus:0.137794,Lama_glama:0.108451)0.900000:0.023382)0.941667:0.021488)0.552778:0.043837,(Equus_caballus:0.084653,Mesohippus_bairdi:0.217741)0.727778:0.047298)0.541667:0.033649)0.888889:0.090993)0.447222:0.050686)0.441667:0.046499)0.250000:0.038994)0.280556:0.052584)0.230556:0.032834,(PA5:0.002810,((((Tribosphenomys_minutus:0.113349,(((Oryctolagus_cuniculus:0.095292,Dawsonolagus_antiquus:0.025286)0.975000:0.033351,Gomphos_elkema:0.080062)1.000000:0.061335,Rhombomylus_turpanensis:0.085632)0.647222:0.021072)0.769444:0.029875,((((PA28:0.000001,(Castor_canadensis:0.090868,Cavia_porcellus:0.087852)0.722222:0.012953)0.863889:0.035560,Ictidomys_tridecemlineatus:0.073231)0.813889:0.041564,Rattus_norvegicus:0.132628)0.911111:0.037429,(Paramys_delicatus:0.052513,Cocomys_lingchaensis:0.072079)0.755556:0.029573)0.669444:0.024781)1.000000:0.099449,(Leptictis_dakotensis:0.180101,(((Moeritherium_lyonsi:0.184286,Carodnia_vieirai:0.092805)0.308333:0.046640,Thomashuxleya_externa_comb:0.136022)0.344444:0.058472,((Phenacodus_intermedius:0.089324,Hyopsodus_paulus:0.101932)0.463889:0.027026,((Protungulatum_donnae:0.136006,Apheliscus_insidiosus:0.101651)0.194444:0.033675,(Didolodus_multicuspis:0.074958,Protolipterna_ellipsodontoides:0.048804)0.583333:0.028159)0.072222:0.018149)0.494444:0.036346)0.213889:0.033748)0.127778:0.040244)0.097222:0.035541,(((((Amblysomus_hottentotus:0.239214,PA44:0.000001)0.838889:0.031142,Echinops_telfairi:0.114842)0.536111:0.031782,PA43:0.010739)0.444444:0.027175,Rhynchocyon_cirnei:0.095474)0.986111:0.049503,((Orycteropus_afer:0.219214,((Loxodonta_africana:0.191896,((PA41:0.000001,(Procavia_capensis:0.000001,PA42:0.000001)1.000000:0.106102)0.744444:0.040582,Trichechus_manatus:0.186426)0.408333:0.040338)0.408333:0.112239,PA40:0.001713)0.352778:0.018655)0.194444:0.014362,(((PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.159950,(Hapalops_elongatus:0.182001,Tamandua_tetradactyla:0.115701)1.000000:0.082413)0.888889:0.084756,PA38:0.000001)0.600000:0.024823)0.177778:0.027488)0.230556:0.074663)0.163889:0.040028)0.127778:0.020553)0.666667:0.044040,PA3:0.005184)0.666667:0.042068)0.486111:0.041235,((Pucadelphys_andinus:0.124447,Didelphis_virginiana:0.040996)1.000000:0.054770,(Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.063651)1.000000:0.062890)0.505556:0.066978)0.997222:0.079079,(Zhangheotherium_quinquecuspidens:0.059751,Henkelotherium_guimarotae:0.066730)0.950000:0.043470)1.000000:0.114976,Haldanodon_exspectatus:0.090908)1.000000:0.080820,Morganucodon_watsoni:0.008454,Morganucodon_oehleri:0.007686);

["max preservation" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: nodal values = "transfer bootstrap expectation" values]
(((((Eomaia_scansoria:0.050363,(Maelestes_gobiensis:0.094181,(Zalambdalestes_lechei:0.130308,Ukhaatherium_nessovi:0.098018)0.830556:0.061090)0.743056:0.039351)0.876852:0.042127,(((PA1:0.000001,(Ornithorhynchus_anatinus:0.081211,Tachyglossus_aculeatus:0.084164)1.000000:0.137421)0.777778:0.053505,((((PA6:0.000001,((Ptilocercus_lowii:0.065217,(PA27:0.005678,Tupaia_glis:0.048324)0.994444:0.023286)1.000000:0.055449,((((PA33:0.000001,((Notharctus_tenebrosus:0.117803,Tarsius_syrichta:0.129514)0.669444:0.044953,Lemur_catta:0.067176)0.790278:0.025893)0.758333:0.004696,(Saimiri_sciureus:0.066634,Homo_sapiens:0.157486)1.000000:0.055836)1.000000:0.057510,(Galeopterus_variegatus:0.018979,Cynocephalus_volans:0.016461)1.000000:0.143761)0.998413:0.059195,PA26:0.000001)0.983333:0.030960)1.000000:0.095500)0.980787:0.025985,(((((Sorex_araneus:0.131111,Talpa_europaea:0.149889)1.000000:0.072061,Erinaceus_europaeus:0.156707)0.876389:0.037278,Solenodon_paradoxus:0.173395)0.933333:0.045872,PA7:0.000001)0.843056:0.037614,(((Metacheiromys_marshi:0.136425,Manis_pentadactyla:0.180535)0.991667:0.107220,(((Felis_silvestris:0.032719,PA21:0.000001)1.000000:0.045998,Canis_lupus:0.059355)0.986111:0.048479,(Sinopa_rapax:0.109679,(Vulpavus_profectus:0.026275,Vulpavus_ovatus:0.016773)1.000000:0.051105)0.980556:0.065954)0.986667:0.050752)0.757143:0.017507,(PA8:0.009764,((Pteropus_giganteus:0.177525,((Icaronycteris_index:0.025653,Onychonycteris_finneyi:0.031331)1.000000:0.094601,((Pteronotus_parnellii:0.074717,Myotis_lucifugus:0.061565)0.780556:0.026429,(Nycteris_thebaica:0.094885,(Saccopteryx_bilineata:0.051306,Rhinopoma_hardwickii:0.050841)0.733333:0.022187)0.998611:0.029776)0.984722:0.030646)0.993056:0.040086)1.000000:0.104906,(PA9:0.007741,(((Sus_scrofa:0.151556,(((Mesonyx_obtusidens:0.149841,((Basilosaurus_cetoides:0.154393,(Artiocetus_clavis:0.108897,(Caperea_marginata:0.162523,Tursiops_truncatus:0.158446)1.000000:0.100905)0.712500:0.039164)0.981481:0.057662,Rodhocetus_balochistanensis:0.099855)0.921528:0.036812)0.890556:0.045237,Archaeotherium_mortoni:0.118788)0.929167:0.056904,Hippopotamus_amphibius:0.152424)0.901984:0.060410)0.865972:0.041576,(PA10:0.000001,(Bos_taurus:0.137794,Lama_glama:0.108451)0.900000:0.023382)0.968056:0.021488)0.880051:0.043837,(Equus_caballus:0.084653,Mesohippus_bairdi:0.217741)0.727778:0.047298)0.934188:0.033649)0.963294:0.090993)0.818182:0.050686)0.819203:0.046499)0.805108:0.038994)0.843056:0.052584)0.825227:0.032834,(PA5:0.002810,((((Tribosphenomys_minutus:0.113349,(((Oryctolagus_cuniculus:0.095292,Dawsonolagus_antiquus:0.025286)0.975000:0.033351,Gomphos_elkema:0.080062)1.000000:0.061335,Rhombomylus_turpanensis:0.085632)0.882407:0.021072)0.940278:0.029875,((((PA28:0.000001,(Castor_canadensis:0.090868,Cavia_porcellus:0.087852)0.722222:0.012953)0.930556:0.035560,Ictidomys_tridecemlineatus:0.073231)0.937037:0.041564,Rattus_norvegicus:0.132628)0.972917:0.037429,(Paramys_delicatus:0.052513,Cocomys_lingchaensis:0.072079)0.755556:0.029573)0.894907:0.024781)1.000000:0.099449,(Leptictis_dakotensis:0.180101,(((Moeritherium_lyonsi:0.184286,Carodnia_vieirai:0.092805)0.308333:0.046640,Thomashuxleya_externa_comb:0.136022)0.629167:0.058472,((Phenacodus_intermedius:0.089324,Hyopsodus_paulus:0.101932)0.463889:0.027026,((Protungulatum_donnae:0.136006,Apheliscus_insidiosus:0.101651)0.194444:0.033675,(Didolodus_multicuspis:0.074958,Protolipterna_ellipsodontoides:0.048804)0.583333:0.028159)0.483333:0.018149)0.833333:0.036346)0.837847:0.033748)0.801543:0.040244)0.732011:0.035541,(((((Amblysomus_hottentotus:0.239214,PA44:0.000001)0.838889:0.031142,Echinops_telfairi:0.114842)0.766667:0.031782,PA43:0.010739)0.814815:0.027175,Rhynchocyon_cirnei:0.095474)0.996528:0.049503,((Orycteropus_afer:0.219214,((Loxodonta_africana:0.191896,((PA41:0.000001,(Procavia_capensis:0.000001,PA42:0.000001)1.000000:0.106102)0.872222:0.040582,Trichechus_manatus:0.186426)0.798148:0.040338)0.834028:0.112239,PA40:0.001713)0.833333:0.018655)0.767130:0.014362,(((PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.159950,(Hapalops_elongatus:0.182001,Tamandua_tetradactyla:0.115701)1.000000:0.082413)0.925926:0.084756,PA38:0.000001)0.869444:0.024823)0.730556:0.027488)0.900521:0.074663)0.736038:0.040028)0.733761:0.020553)0.922049:0.044040,PA3:0.005184)0.917407:0.042068)0.778009:0.041235,((Pucadelphys_andinus:0.124447,Didelphis_virginiana:0.040996)1.000000:0.054770,(Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.063651)1.000000:0.062890)0.762500:0.066978)0.998611:0.079079,(Zhangheotherium_quinquecuspidens:0.059751,Henkelotherium_guimarotae:0.066730)0.950000:0.043470)1.000000:0.114976,Haldanodon_exspectatus:0.090908)1.000000:0.080820,Morganucodon_watsoni:0.008454,Morganucodon_oehleri:0.007686);

["max preservation" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: nodal values = standard bootstrap values]
(Morganucodon_watsoni:0.008521,((((Eomaia_scansoria:0.050685,(Maelestes_gobiensis:0.094747,(Ukhaatherium_nessovi:0.099039,Zalambdalestes_lechei:0.131062)0.786765:0.061636)0.470588:0.039830)0.708333:0.042815,((((((PA6:0.000956,((Galeopterus_variegatus:0.020438,Cynocephalus_volans:0.015243)1.000000:0.171785,((Tupaia_glis:0.061859,Ptilocercus_lowii:0.075473)1.000000:0.053244,(((Tarsius_syrichta:0.144658,(Homo_sapiens:0.155372,Saimiri_sciureus:0.072888)1.000000:0.049663)0.698529:0.023649,(Notharctus_tenebrosus:0.132202,Lemur_catta:0.062764)0.843137:0.039889)1.000000:0.072158,PA26:0.001759)0.649510:0.017610)0.968137:0.032801)1.000000:0.076841)0.936275:0.032721,((Solenodon_paradoxus:0.178377,((Sorex_araneus:0.146661,(Talpa_europaea:0.128561,Amblysomus_hottentotus:0.240172)0.406863:0.046262)0.757353:0.059678,(Echinops_telfairi:0.161253,Erinaceus_europaeus:0.115040)0.691176:0.054811)0.514706:0.029580)0.419118:0.036135,((((Felis_silvestris:0.088663,Canis_lupus:0.073600)0.987745:0.044283,((Vulpavus_ovatus:0.018398,Vulpavus_profectus:0.025312)1.000000:0.055608,Sinopa_rapax:0.107104)0.436275:0.040466)1.000000:0.073493,(Manis_pentadactyla:0.191864,Metacheiromys_marshi:0.127690)0.997549:0.105579)0.178922:0.033007,(PA7:0.011254,((((Myotis_lucifugus:0.062310,Pteronotus_parnellii:0.074573)0.752451:0.025924,(Nycteris_thebaica:0.095261,(Saccopteryx_bilineata:0.052599,Rhinopoma_hardwickii:0.051027)0.742647:0.023549)1.000000:0.031593)0.995098:0.038563,(Icaronycteris_index:0.025530,Onychonycteris_finneyi:0.031691)1.000000:0.082663)0.968137:0.040746,Pteropus_giganteus:0.178344)1.000000:0.147716)0.455882:0.018027)0.107843:0.030486)0.080882:0.031100)0.078431:0.028549,(PA5:0.007341,((Leptictis_dakotensis:0.215715,(Rhynchocyon_cirnei:0.128727,((Orycteropus_afer:0.224025,(((Loxodonta_africana:0.183864,(Moeritherium_lyonsi:0.152075,Trichechus_manatus:0.162402)0.294118:0.046388)0.661765:0.087884,Procavia_capensis:0.185624)0.193627:0.045246,PA40:0.002830)0.193627:0.018580)0.112745:0.015646,(PA38:0.000001,((Tamandua_tetradactyla:0.118089,Hapalops_elongatus:0.184851)1.000000:0.083603,(PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.163429)0.835784:0.086804)0.477941:0.022569)0.095588:0.038204)0.102941:0.048955)0.017157:0.027950,((((Cocomys_lingchaensis:0.074978,Paramys_delicatus:0.051440)0.409314:0.028103,(((Castor_canadensis:0.084766,Cavia_porcellus:0.091491)0.948529:0.047127,Ictidomys_tridecemlineatus:0.067084)0.509804:0.044013,Rattus_norvegicus:0.148928)0.316176:0.024165)0.651961:0.030501,(Tribosphenomys_minutus:0.114736,(Rhombomylus_turpanensis:0.077429,((Dawsonolagus_antiquus:0.025332,Oryctolagus_cuniculus:0.099371)0.973039:0.033149,Gomphos_elkema:0.078790)1.000000:0.071024)0.622549:0.022102)0.838235:0.034153)1.000000:0.091777,((Mesonyx_obtusidens:0.172545,((((Sus_scrofa:0.175418,(((Artiocetus_clavis:0.134532,((Tursiops_truncatus:0.169296,Caperea_marginata:0.159938)0.997549:0.109212,Basilosaurus_cetoides:0.135800)0.578431:0.047122)0.953431:0.107199,Archaeotherium_mortoni:0.110632)0.169118:0.036428,Hippopotamus_amphibius:0.173883)0.137255:0.042417)0.068627:0.030786,(Lama_glama:0.119490,Bos_taurus:0.126345)0.997549:0.056755)0.073529:0.029554,(Mesohippus_bairdi:0.153183,Equus_caballus:0.143014)0.970588:0.060671)0.105392:0.026670,Rodhocetus_balochistanensis:0.141986)0.232843:0.018558)0.901961:0.044737,(((Hyopsodus_paulus:0.100535,Phenacodus_intermedius:0.092136)0.458333:0.023474,((Apheliscus_insidiosus:0.101724,Protungulatum_donnae:0.139926)0.186275:0.036262,(Protolipterna_ellipsodontoides:0.054513,Didolodus_multicuspis:0.069577)0.583333:0.026974)0.056373:0.018484)0.575980:0.037092,(Thomashuxleya_externa_comb:0.127867,Carodnia_vieirai:0.120244)0.259804:0.055802)0.120098:0.040821)0.129902:0.039379)0.098039:0.038833)0.078431:0.033593)0.075980:0.018795)0.634804:0.041544,PA3:0.005937)0.639706:0.042461,((Ornithorhynchus_anatinus:0.081746,Tachyglossus_aculeatus:0.085570)1.000000:0.138585,PA1:0.000001)0.507353:0.052958)0.460784:0.041445,((Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.064286,(Didelphis_virginiana:0.041324,Pucadelphys_andinus:0.125411)1.000000:0.055302)1.000000:0.064455)0.475490:0.066668)0.997549:0.079344,(Henkelotherium_guimarotae:0.067260,Zhangheotherium_quinquecuspidens:0.060085)0.933824:0.043744)1.000000:0.115689,Haldanodon_exspectatus:0.091542)1.000000:0.081712,Morganucodon_oehleri:0.007790);

["max preservation" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: nodal values = "transfer bootstrap expectation" values]
(Morganucodon_watsoni:0.008521,((((Eomaia_scansoria:0.050685,(Maelestes_gobiensis:0.094747,(Ukhaatherium_nessovi:0.099039,Zalambdalestes_lechei:0.131062)0.786765:0.061636)0.732843:0.039830)0.892974:0.042815,((((((PA6:0.000956,((Galeopterus_variegatus:0.020438,Cynocephalus_volans:0.015243)1.000000:0.171785,((Tupaia_glis:0.061859,Ptilocercus_lowii:0.075473)1.000000:0.053244,(((Tarsius_syrichta:0.144658,(Homo_sapiens:0.155372,Saimiri_sciureus:0.072888)1.000000:0.049663)0.849265:0.023649,(Notharctus_tenebrosus:0.132202,Lemur_catta:0.062764)0.843137:0.039889)1.000000:0.072158,PA26:0.001759)0.929902:0.017610)0.990896:0.032801)1.000000:0.076841)0.993627:0.032721,((Solenodon_paradoxus:0.178377,((Sorex_araneus:0.146661,(Talpa_europaea:0.128561,Amblysomus_hottentotus:0.240172)0.406863:0.046262)0.878676:0.059678,(Echinops_telfairi:0.161253,Erinaceus_europaeus:0.115040)0.691176:0.054811)0.767770:0.029580)0.777451:0.036135,((((Felis_silvestris:0.088663,Canis_lupus:0.073600)0.987745:0.044283,((Vulpavus_ovatus:0.018398,Vulpavus_profectus:0.025312)1.000000:0.055608,Sinopa_rapax:0.107104)0.718137:0.040466)1.000000:0.073493,(Manis_pentadactyla:0.191864,Metacheiromys_marshi:0.127690)0.997549:0.105579)0.727533:0.033007,(PA7:0.011254,((((Myotis_lucifugus:0.062310,Pteronotus_parnellii:0.074573)0.752451:0.025924,(Nycteris_thebaica:0.095261,(Saccopteryx_bilineata:0.052599,Rhinopoma_hardwickii:0.051027)0.742647:0.023549)1.000000:0.031593)0.997549:0.038563,(Icaronycteris_index:0.025530,Onychonycteris_finneyi:0.031691)1.000000:0.082663)0.994690:0.040746,Pteropus_giganteus:0.178344)1.000000:0.147716)0.931985:0.018027)0.716667:0.030486)0.765873:0.031100)0.810355:0.028549,(PA5:0.007341,((Leptictis_dakotensis:0.215715,(Rhynchocyon_cirnei:0.128727,((Orycteropus_afer:0.224025,(((Loxodonta_africana:0.183864,(Moeritherium_lyonsi:0.152075,Trichechus_manatus:0.162402)0.294118:0.046388)0.825980:0.087884,Procavia_capensis:0.185624)0.667484:0.045246,PA40:0.002830)0.591299:0.018580)0.528922:0.015646,(PA38:0.000001,((Tamandua_tetradactyla:0.118089,Hapalops_elongatus:0.184851)1.000000:0.083603,(PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.163429)0.890523:0.086804)0.815564:0.022569)0.643627:0.038204)0.716355:0.048955)0.663194:0.027950,((((Cocomys_lingchaensis:0.074978,Paramys_delicatus:0.051440)0.409314:0.028103,(((Castor_canadensis:0.084766,Cavia_porcellus:0.091491)0.948529:0.047127,Ictidomys_tridecemlineatus:0.067084)0.754902:0.044013,Rattus_norvegicus:0.148928)0.671569:0.024165)0.866176:0.030501,(Tribosphenomys_minutus:0.114736,(Rhombomylus_turpanensis:0.077429,((Dawsonolagus_antiquus:0.025332,Oryctolagus_cuniculus:0.099371)0.973039:0.033149,Gomphos_elkema:0.078790)1.000000:0.071024)0.874183:0.022102)0.952819:0.034153)1.000000:0.091777,((Mesonyx_obtusidens:0.172545,((((Sus_scrofa:0.175418,(((Artiocetus_clavis:0.134532,((Tursiops_truncatus:0.169296,Caperea_marginata:0.159938)0.997549:0.109212,Basilosaurus_cetoides:0.135800)0.789216:0.047122)0.981209:0.107199,Archaeotherium_mortoni:0.110632)0.780637:0.036428,Hippopotamus_amphibius:0.173883)0.691176:0.042417)0.611111:0.030786,(Lama_glama:0.119490,Bos_taurus:0.126345)0.997549:0.056755)0.611520:0.029554,(Mesohippus_bairdi:0.153183,Equus_caballus:0.143014)0.970588:0.060671)0.823284:0.026670,Rodhocetus_balochistanensis:0.141986)0.900847:0.018558)0.971609:0.044737,(((Hyopsodus_paulus:0.100535,Phenacodus_intermedius:0.092136)0.458333:0.023474,((Apheliscus_insidiosus:0.101724,Protungulatum_donnae:0.139926)0.186275:0.036262,(Protolipterna_ellipsodontoides:0.054513,Didolodus_multicuspis:0.069577)0.583333:0.026974)0.457516:0.018484)0.888235:0.037092,(Thomashuxleya_externa_comb:0.127867,Carodnia_vieirai:0.120244)0.259804:0.055802)0.745448:0.040821)0.815196:0.039379)0.819892:0.038833)0.847148:0.033593)0.845316:0.018795)0.912071:0.041544,PA3:0.005937)0.907190:0.042461,((Ornithorhynchus_anatinus:0.081746,Tachyglossus_aculeatus:0.085570)1.000000:0.138585,PA1:0.000001)0.753676:0.052958)0.758374:0.041445,((Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.064286,(Didelphis_virginiana:0.041324,Pucadelphys_andinus:0.125411)1.000000:0.055302)1.000000:0.064455)0.745404:0.066668)0.998775:0.079344,(Henkelotherium_guimarotae:0.067260,Zhangheotherium_quinquecuspidens:0.060085)0.933824:0.043744)1.000000:0.115689,Haldanodon_exspectatus:0.091542)1.000000:0.081712,Morganucodon_oehleri:0.007790);

["typical preservation" PAs corrected for the O'Leary et al. character ontology: nodal values = standard bootstrap values]
((Haldanodon_exspectatus:0.097264,((Henkelotherium_guimarotae:0.077990,Zhangheotherium_quinquecuspidens:0.054411)0.956667:0.055320,((Tachyglossus_aculeatus:0.087206,(Ornithorhynchus_anatinus:0.000001,PA2:0.000001)1.000000:0.072375)1.000000:0.105665,(PA1:0.000001,((PA3:0.002775,((((((((PA41:0.000001,(PA42:0.000001,Procavia_capensis:0.000001)1.000000:0.111392)0.656667:0.065183,((Loxodonta_africana:0.151112,Moeritherium_lyonsi:0.171614)0.276667:0.056863,Trichechus_manatus:0.185920)0.186667:0.034746)0.456667:0.108129,PA40:0.001936)0.293333:0.021601,((((Manis_pentadactyla:0.187152,Metacheiromys_marshi:0.117777)0.996667:0.085733,((Tamandua_tetradactyla:0.137739,Hapalops_elongatus:0.164183)1.000000:0.075798,(Dasypus_novemcinctus:0.000001,PA39:0.000001)1.000000:0.171568)0.746667:0.041306)0.833333:0.046466,Orycteropus_afer:0.219930)0.300000:0.029231,PA38:0.004200)0.370000:0.016211)0.153333:0.029031,(Rhynchocyon_cirnei:0.123118,(((Echinops_telfairi:0.000001,PA45:0.000001)1.000000:0.127572,(Amblysomus_hottentotus:0.231723,PA44:0.000693)0.920000:0.031294)0.863333:0.032884,PA43:0.013755)0.833333:0.031525)0.993333:0.040967)0.270000:0.064228,(((Rhombomylus_turpanensis:0.083929,((Gomphos_elkema:0.075955,(Dawsonolagus_antiquus:0.025548,Oryctolagus_cuniculus:0.096492)0.986667:0.038983)1.000000:0.059050,Tribosphenomys_minutus:0.111274)0.203333:0.015872)0.836667:0.044605,((Cocomys_lingchaensis:0.073425,Paramys_delicatus:0.051711)0.783333:0.027353,(((Ictidomys_tridecemlineatus:0.000001,PA31:0.000001)1.000000:0.064259,(PA28:0.019252,(((PA30:0.000001,Castor_canadensis:0.000001)1.000000:0.071530,PA29:0.004248)0.976667:0.025143,Cavia_porcellus:0.086693)0.573333:0.014440)0.743333:0.028698)0.880000:0.046169,Rattus_norvegicus:0.138138)0.893333:0.033794)0.763333:0.028718)1.000000:0.099043,(Leptictis_dakotensis:0.175097,(Protungulatum_donnae:0.135299,((((Carodnia_vieirai:0.122120,Thomashuxleya_externa_comb:0.123534)0.510000:0.075291,(Hyopsodus_paulus:0.102741,Phenacodus_intermedius:0.088460)0.450000:0.024666)0.063333:0.015605,(Protolipterna_ellipsodontoides:0.050828,Didolodus_multicuspis:0.072044)0.630000:0.029993)0.090000:0.017206,Apheliscus_insidiosus:0.119744)0.136667:0.019149)0.193333:0.032422)0.050000:0.041672)0.010000:0.028757)0.046667:0.036518,PA5:0.001320)0.033333:0.016605,(((Ptilocercus_lowii:0.072898,(PA27:0.001206,(PA32:0.000001,Tupaia_glis:0.000001)1.000000:0.037201)1.000000:0.027254)1.000000:0.061797,(PA26:0.000001,((PA33:0.005372,((((PA36:0.005674,(Saimiri_sciureus:0.000001,PA37:0.000001)1.000000:0.051861)0.486667:0.010641,Homo_sapiens:0.154693)1.000000:0.055847,(Tarsius_syrichta:0.128575,PA35:0.015047)0.313333:0.014108)0.580000:0.033983,(Notharctus_tenebrosus:0.139448,Lemur_catta:0.055886)0.710000:0.030391)0.690000:0.014304)1.000000:0.058768,(Galeopterus_variegatus:0.018772,(Cynocephalus_volans:0.000001,PA34:0.000001)1.000000:0.016729)1.000000:0.156970)0.786667:0.042156)0.573333:0.024255)1.000000:0.079314,(PA6:0.000164,(((PA8:0.010382,((Mesonyx_obtusidens:0.151964,(((Artiocetus_clavis:0.123815,(Basilosaurus_cetoides:0.134540,(Caperea_marginata:0.148782,(PA15:0.000001,Tursiops_truncatus:0.000001)1.000000:0.152530)1.000000:0.126275)0.610000:0.046568)0.936667:0.047079,Rodhocetus_balochistanensis:0.103023)0.603333:0.049646,Archaeotherium_mortoni:0.135339)0.273333:0.032201)0.826667:0.085173,(PA9:0.025366,((PA10:0.012383,(Lama_glama:0.101220,(((PA13:0.012648,Bos_taurus:0.035368)0.996667:0.065212,((Hippopotamus_amphibius:0.000001,PA14:0.000001)1.000000:0.196798,(PA12:0.000001,Sus_scrofa:0.155600)0.603333:0.011898)0.530000:0.039245)0.696667:0.040434,PA11:0.005181)0.613333:0.028702)0.686667:0.029763)0.740000:0.040697,(Mesohippus_bairdi:0.203384,Equus_caballus:0.102943)0.533333:0.053346)0.263333:0.023223)0.646667:0.042696)0.646667:0.056279)0.376667:0.045640,((Canis_lupus:0.061854,(PA21:0.000001,(Felis_silvestris:0.000001,PA22:0.000001)1.000000:0.028122)1.000000:0.051316)0.993333:0.049049,(Sinopa_rapax:0.108011,(Vulpavus_profectus:0.027767,Vulpavus_ovatus:0.015746)1.000000:0.052567)0.543333:0.061306)0.996667:0.076912)0.056667:0.041143,((Pteropus_giganteus:0.170366,((Onychonycteris_finneyi:0.032611,Icaronycteris_index:0.024073)1.000000:0.101134,((Rhinopoma_hardwickii:0.000001,PA20:0.000001)1.000000:0.052766,((((PA18:0.006442,Nycteris_thebaica:0.090451)0.930000:0.011197,(Saccopteryx_bilineata:0.000001,PA19:0.000001)1.000000:0.047516)1.000000:0.027485,(Pteronotus_parnellii:0.072783,(PA17:0.000001,Myotis_lucifugus:0.053096)0.960000:0.009509)0.863333:0.024834)0.796667:0.024712,PA16:0.000001)0.706667:0.014067)0.933333:0.052494)0.653333:0.028680)1.000000:0.141448,((Solenodon_paradoxus:0.126239,(PA23:0.000001,(Erinaceus_europaeus:0.098320,(PA24:0.000001,(Talpa_europaea:0.124258,(Sorex_araneus:0.000001,PA25:0.000001)1.000000:0.104491)0.990000:0.071894)0.640000:0.038682)0.873333:0.053473)0.810000:0.045419)1.000000:0.083092,PA7:0.003448)0.443333:0.018690)0.153333:0.028069)0.103333:0.044457)0.073333:0.020502)0.066667:0.025638)0.643333:0.048153)0.666667:0.030539,(((Pucadelphys_andinus:0.119712,Didelphis_virginiana:0.049006)1.000000:0.059858,(Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.073732)1.000000:0.051012,((Zalambdalestes_lechei:0.123511,Ukhaatherium_nessovi:0.106386)0.840000:0.059127,(Maelestes_gobiensis:0.094981,Eomaia_scansoria:0.066013)0.623333:0.034478)0.823333:0.083096)0.260000:0.041740)0.506667:0.053774)0.790000:0.076356)0.706667:0.080206)1.000000:0.100977)1.000000:0.077630,Morganucodon_watsoni:0.007960,Morganucodon_oehleri:0.008535);

["typical preservation" PAs corrected for the O'Leary et al. character ontology: nodal values = "transfer bootstrap expectation" values]
((Haldanodon_exspectatus:0.097264,((Henkelotherium_guimarotae:0.077990,Zhangheotherium_quinquecuspidens:0.054411)0.956667:0.055320,((Tachyglossus_aculeatus:0.087206,(Ornithorhynchus_anatinus:0.000001,PA2:0.000001)1.000000:0.072375)1.000000:0.105665,(PA1:0.000001,((PA3:0.002775,((((((((PA41:0.000001,(PA42:0.000001,Procavia_capensis:0.000001)1.000000:0.111392)0.828333:0.065183,((Loxodonta_africana:0.151112,Moeritherium_lyonsi:0.171614)0.276667:0.056863,Trichechus_manatus:0.185920)0.341667:0.034746)0.886000:0.108129,PA40:0.001936)0.861111:0.021601,((((Manis_pentadactyla:0.187152,Metacheiromys_marshi:0.117777)0.996667:0.085733,((Tamandua_tetradactyla:0.137739,Hapalops_elongatus:0.164183)1.000000:0.075798,(Dasypus_novemcinctus:0.000001,PA39:0.000001)1.000000:0.171568)0.831111:0.041306)0.933333:0.046466,Orycteropus_afer:0.219930)0.830556:0.029231,PA38:0.004200)0.831429:0.016211)0.721429:0.029031,(Rhynchocyon_cirnei:0.123118,(((Echinops_telfairi:0.000001,PA45:0.000001)1.000000:0.127572,(Amblysomus_hottentotus:0.231723,PA44:0.000693)0.920000:0.031294)0.920000:0.032884,PA43:0.013755)0.958333:0.031525)0.998667:0.040967)0.893667:0.064228,(((Rhombomylus_turpanensis:0.083929,((Gomphos_elkema:0.075955,(Dawsonolagus_antiquus:0.025548,Oryctolagus_cuniculus:0.096492)0.986667:0.038983)1.000000:0.059050,Tribosphenomys_minutus:0.111274)0.734444:0.015872)0.958333:0.044605,((Cocomys_lingchaensis:0.073425,Paramys_delicatus:0.051711)0.783333:0.027353,(((Ictidomys_tridecemlineatus:0.000001,PA31:0.000001)1.000000:0.064259,(PA28:0.019252,(((PA30:0.000001,Castor_canadensis:0.000001)1.000000:0.071530,PA29:0.004248)0.988333:0.025143,Cavia_porcellus:0.086693)0.851111:0.014440)0.872500:0.028698)0.977778:0.046169,Rattus_norvegicus:0.138138)0.982381:0.033794)0.949259:0.028718)1.000000:0.099043,(Leptictis_dakotensis:0.175097,(Protungulatum_donnae:0.135299,((((Carodnia_vieirai:0.122120,Thomashuxleya_externa_comb:0.123534)0.510000:0.075291,(Hyopsodus_paulus:0.102741,Phenacodus_intermedius:0.088460)0.450000:0.024666)0.340000:0.015605,(Protolipterna_ellipsodontoides:0.050828,Didolodus_multicuspis:0.072044)0.630000:0.029993)0.578000:0.017206,Apheliscus_insidiosus:0.119744)0.722222:0.019149)0.767143:0.032422)0.698333:0.041672)0.652899:0.028757)0.623864:0.036518,PA5:0.001320)0.623037:0.016605,(((Ptilocercus_lowii:0.072898,(PA27:0.001206,(PA32:0.000001,Tupaia_glis:0.000001)1.000000:0.037201)1.000000:0.027254)1.000000:0.061797,(PA26:0.000001,((PA33:0.005372,((((PA36:0.005674,(Saimiri_sciureus:0.000001,PA37:0.000001)1.000000:0.051861)0.743333:0.010641,Homo_sapiens:0.154693)1.000000:0.055847,(Tarsius_syrichta:0.128575,PA35:0.015047)0.313333:0.014108)0.838000:0.033983,(Notharctus_tenebrosus:0.139448,Lemur_catta:0.055886)0.710000:0.030391)0.955714:0.014304)1.000000:0.058768,(Galeopterus_variegatus:0.018772,(Cynocephalus_volans:0.000001,PA34:0.000001)1.000000:0.016729)1.000000:0.156970)0.942424:0.042156)0.915000:0.024255)1.000000:0.079314,(PA6:0.000164,(((PA8:0.010382,((Mesonyx_obtusidens:0.151964,(((Artiocetus_clavis:0.123815,(Basilosaurus_cetoides:0.134540,(Caperea_marginata:0.148782,(PA15:0.000001,Tursiops_truncatus:0.000001)1.000000:0.152530)1.000000:0.126275)0.870000:0.046568)0.981667:0.047079,Rodhocetus_balochistanensis:0.103023)0.916000:0.049646,Archaeotherium_mortoni:0.135339)0.866667:0.032201)0.953810:0.085173,(PA9:0.025366,((PA10:0.012383,(Lama_glama:0.101220,(((PA13:0.012648,Bos_taurus:0.035368)0.996667:0.065212,((Hippopotamus_amphibius:0.000001,PA14:0.000001)1.000000:0.196798,(PA12:0.000001,Sus_scrofa:0.155600)0.603333:0.011898)0.694444:0.039245)0.802667:0.040434,PA11:0.005181)0.802778:0.028702)0.855238:0.029763)0.880000:0.040697,(Mesohippus_bairdi:0.203384,Equus_caballus:0.102943)0.533333:0.053346)0.853667:0.023223)0.907273:0.042696)0.876491:0.056279)0.868000:0.045640,((Canis_lupus:0.061854,(PA21:0.000001,(Felis_silvestris:0.000001,PA22:0.000001)1.000000:0.028122)1.000000:0.051316)0.997778:0.049049,(Sinopa_rapax:0.108011,(Vulpavus_profectus:0.027767,Vulpavus_ovatus:0.015746)1.000000:0.052567)0.771667:0.061306)0.999444:0.076912)0.668025:0.041143,((Pteropus_giganteus:0.170366,((Onychonycteris_finneyi:0.032611,Icaronycteris_index:0.024073)1.000000:0.101134,((Rhinopoma_hardwickii:0.000001,PA20:0.000001)1.000000:0.052766,((((PA18:0.006442,Nycteris_thebaica:0.090451)0.930000:0.011197,(Saccopteryx_bilineata:0.000001,PA19:0.000001)1.000000:0.047516)1.000000:0.027485,(Pteronotus_parnellii:0.072783,(PA17:0.000001,Myotis_lucifugus:0.053096)0.960000:0.009509)0.926667:0.024834)0.909444:0.024712,PA16:0.000001)0.928571:0.014067)0.985185:0.052494)0.968485:0.028680)1.000000:0.141448,((Solenodon_paradoxus:0.126239,(PA23:0.000001,(Erinaceus_europaeus:0.098320,(PA24:0.000001,(Talpa_europaea:0.124258,(Sorex_araneus:0.000001,PA25:0.000001)1.000000:0.104491)0.995000:0.071894)0.880000:0.038682)0.965833:0.053473)0.962000:0.045419)1.000000:0.083092,PA7:0.003448)0.920476:0.018690)0.714667:0.028069)0.689375:0.044457)0.687279:0.020502)0.653280:0.025638)0.929608:0.048153)0.926042:0.030539,(((Pucadelphys_andinus:0.119712,Didelphis_virginiana:0.049006)1.000000:0.059858,(Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.073732)1.000000:0.051012,((Zalambdalestes_lechei:0.123511,Ukhaatherium_nessovi:0.106386)0.840000:0.059127,(Maelestes_gobiensis:0.094981,Eomaia_scansoria:0.066013)0.623333:0.034478)0.936667:0.083096)0.581429:0.041740)0.870833:0.053774)0.914762:0.076356)0.854167:0.080206)1.000000:0.100977)1.000000:0.077630,Morganucodon_watsoni:0.007960,Morganucodon_oehleri:0.008535);

["typical preservation" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: nodal values = standard bootstrap values]
((Haldanodon_exspectatus:0.097557,((Zhangheotherium_quinquecuspidens:0.054350,Henkelotherium_guimarotae:0.078203)0.929348:0.055604,((Ornithorhynchus_anatinus:0.073416,Tachyglossus_aculeatus:0.088025)1.000000:0.102685,(PA1:0.000001,((((Zalambdalestes_lechei:0.123757,Ukhaatherium_nessovi:0.106441)0.831522:0.059507,(Maelestes_gobiensis:0.095441,Eomaia_scansoria:0.066103)0.543478:0.034243)0.811594:0.082159,((Pucadelphys_andinus:0.119690,Didelphis_virginiana:0.049331)0.990942:0.059558,(PA4:0.000001,Dromiciops_gliroides:0.000001)1.000000:0.073423)1.000000:0.051428)0.260870:0.041413,(((PA6:0.022586,(((PA8:0.031189,(PA9:0.019926,((Mesohippus_bairdi:0.201921,Equus_caballus:0.102360)0.672101:0.048854,(((((Mesonyx_obtusidens:0.149207,(Rodhocetus_balochistanensis:0.096813,((Basilosaurus_cetoides:0.140977,(Caperea_marginata:0.166489,Tursiops_truncatus:0.163104)1.000000:0.107785)0.579710:0.044256,Artiocetus_clavis:0.123473)0.940217:0.047454)0.628623:0.043269)0.398551:0.044660,Archaeotherium_mortoni:0.118291)0.577899:0.052069,Hippopotamus_amphibius:0.160307)0.423913:0.057393,Sus_scrofa:0.156647)0.351449:0.040785,(PA10:0.000208,(Lama_glama:0.114620,Bos_taurus:0.132774)0.826087:0.023328)0.978261:0.025650)0.338768:0.037286)0.297101:0.027205)0.885870:0.083841)0.436594:0.035250,(Pteropus_giganteus:0.174749,((Icaronycteris_index:0.025050,Onychonycteris_finneyi:0.032278)1.000000:0.086056,((Myotis_lucifugus:0.062088,Pteronotus_parnellii:0.074771)0.820652:0.027495,(Nycteris_thebaica:0.094650,(Saccopteryx_bilineata:0.051189,Rhinopoma_hardwickii:0.052162)0.760870:0.022928)0.996377:0.030232)0.967391:0.036963)0.952899:0.040858)1.000000:0.116749)0.278986:0.036073,(PA7:0.002496,((Erinaceus_europaeus:0.158974,(Sorex_araneus:0.133806,Talpa_europaea:0.148872)1.000000:0.073075)0.617754:0.037754,Solenodon_paradoxus:0.174863)0.592391:0.032631)0.445652:0.031899)0.192029:0.040135)0.086957:0.027383,((((Ptilocercus_lowii:0.066507,(PA27:0.000001,Tupaia_glis:0.036261)1.000000:0.033587)1.000000:0.071689,PA26:0.001202)0.405797:0.022566,((Galeopterus_variegatus:0.018660,Cynocephalus_volans:0.017034)1.000000:0.155014,((PA33:0.000001,((Tarsius_syrichta:0.132321,Notharctus_tenebrosus:0.112512)0.648551:0.042668,Lemur_catta:0.072152)0.557971:0.028125)0.391304:0.003990,(Homo_sapiens:0.154176,Saimiri_sciureus:0.072750)1.000000:0.056539)1.000000:0.060601)0.836957:0.034360)1.000000:0.098444,((((Rhynchocyon_cirnei:0.122845,((Echinops_telfairi:0.117599,(Amblysomus_hottentotus:0.231515,PA44:0.000001)0.923913:0.032342)0.804348:0.032872,PA43:0.013345)0.768116:0.031630)0.985507:0.042398,((PA38:0.004263,(((Manis_pentadactyla:0.184275,Metacheiromys_marshi:0.117192)0.996377:0.084246,((PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.169768,(Hapalops_elongatus:0.162070,Tamandua_tetradactyla:0.137381)0.996377:0.075768)0.755435:0.042032)0.807971:0.045350,Orycteropus_afer:0.220292)0.311594:0.029253)0.342391:0.016334,(((PA42:0.000001,Procavia_capensis:0.000001)1.000000:0.098213,(Loxodonta_africana:0.232250,(Trichechus_manatus:0.207714,PA41:0.000001)0.260870:0.001159)0.204710:0.045642)0.405797:0.133012,PA40:0.002109)0.273551:0.021154)0.152174:0.027786)0.235507:0.065040,(((((Thomashuxleya_externa_comb:0.127444,Carodnia_vieirai:0.108771)0.500000:0.036299,Moeritherium_lyonsi:0.206572)0.286232:0.057975,((Hyopsodus_paulus:0.102966,Phenacodus_intermedius:0.087784)0.452899:0.026999,(Apheliscus_insidiosus:0.118428,(Protolipterna_ellipsodontoides:0.049059,Didolodus_multicuspis:0.076745)0.639493:0.027521)0.163043:0.018872)0.307971:0.029710)0.077899:0.036770,((Protungulatum_donnae:0.127717,Leptictis_dakotensis:0.159458)0.119565:0.026168,(Sinopa_rapax:0.095635,((Vulpavus_ovatus:0.018580,Vulpavus_profectus:0.025011)1.000000:0.064319,(Canis_lupus:0.052167,(PA21:0.000001,Felis_silvestris:0.033050)1.000000:0.059180)0.998188:0.068889)0.409420:0.023900)1.000000:0.071535)0.076087:0.027674)0.018116:0.034984,((Tribosphenomys_minutus:0.110691,(Rhombomylus_turpanensis:0.081121,(Gomphos_elkema:0.078261,(Dawsonolagus_antiquus:0.025357,Oryctolagus_cuniculus:0.099436)0.958333:0.034592)1.000000:0.065339)0.659420:0.022516)0.780797:0.032247,((Cocomys_lingchaensis:0.074266,Paramys_delicatus:0.051439)0.746377:0.025948,((Ictidomys_tridecemlineatus:0.065953,(PA28:0.000001,(Castor_canadensis:0.093053,Cavia_porcellus:0.086496)0.637681:0.013102)0.903986:0.039360)0.804348:0.044737,Rattus_norvegicus:0.139132)0.851449:0.035999)0.711957:0.026291)1.000000:0.097708)0.014493:0.032620)0.048913:0.036437,PA5:0.001318)0.034420:0.011388)0.016304:0.014105)0.693841:0.045230,PA3:0.003055)0.708333:0.029833)0.463768:0.053954)0.807971:0.075931)0.652174:0.080480)1.000000:0.100698)1.000000:0.077088,Morganucodon_watsoni:0.007896,Morganucodon_oehleri:0.008586);

["typical preservation" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes above the ordinal level included: nodal values = "transfer bootstrap expectation" values]
((Haldanodon_exspectatus:0.097557,((Zhangheotherium_quinquecuspidens:0.054350,Henkelotherium_guimarotae:0.078203)0.929348:0.055604,((Ornithorhynchus_anatinus:0.073416,Tachyglossus_aculeatus:0.088025)1.000000:0.102685,(PA1:0.000001,((((Zalambdalestes_lechei:0.123757,Ukhaatherium_nessovi:0.106441)0.831522:0.059507,(Maelestes_gobiensis:0.095441,Eomaia_scansoria:0.066103)0.543478:0.034243)0.931763:0.082159,((Pucadelphys_andinus:0.119690,Didelphis_virginiana:0.049331)0.990942:0.059558,(PA4:0.000001,Dromiciops_gliroides:0.000001)1.000000:0.073423)1.000000:0.051428)0.581263:0.041413,(((PA6:0.022586,(((PA8:0.031189,(PA9:0.019926,((Mesohippus_bairdi:0.201921,Equus_caballus:0.102360)0.672101:0.048854,(((((Mesonyx_obtusidens:0.149207,(Rodhocetus_balochistanensis:0.096813,((Basilosaurus_cetoides:0.140977,(Caperea_marginata:0.166489,Tursiops_truncatus:0.163104)1.000000:0.107785)0.789855:0.044256,Artiocetus_clavis:0.123473)0.975845:0.047454)0.902627:0.043269)0.837681:0.044660,Archaeotherium_mortoni:0.118291)0.865942:0.052069,Hippopotamus_amphibius:0.160307)0.828157:0.057393,Sus_scrofa:0.156647)0.772871:0.040785,(PA10:0.000208,(Lama_glama:0.114620,Bos_taurus:0.132774)0.826087:0.023328)0.988225:0.025650)0.818841:0.037286)0.924470:0.027205)0.977355:0.083841)0.947947:0.035250,(Pteropus_giganteus:0.174749,((Icaronycteris_index:0.025050,Onychonycteris_finneyi:0.032278)1.000000:0.086056,((Myotis_lucifugus:0.062088,Pteronotus_parnellii:0.074771)0.820652:0.027495,(Nycteris_thebaica:0.094650,(Saccopteryx_bilineata:0.051189,Rhinopoma_hardwickii:0.052162)0.760870:0.022928)0.998188:0.030232)0.983696:0.036963)0.992150:0.040858)1.000000:0.116749)0.729836:0.036073,(PA7:0.002496,((Erinaceus_europaeus:0.158974,(Sorex_araneus:0.133806,Talpa_europaea:0.148872)1.000000:0.073075)0.808877:0.037754,Solenodon_paradoxus:0.174863)0.840580:0.032631)0.783514:0.031899)0.664402:0.040135)0.647739:0.027383,((((Ptilocercus_lowii:0.066507,(PA27:0.000001,Tupaia_glis:0.036261)1.000000:0.033587)1.000000:0.071689,PA26:0.001202)0.801932:0.022566,((Galeopterus_variegatus:0.018660,Cynocephalus_volans:0.017034)1.000000:0.155014,((PA33:0.000001,((Tarsius_syrichta:0.132321,Notharctus_tenebrosus:0.112512)0.648551:0.042668,Lemur_catta:0.072152)0.776268:0.028125)0.697464:0.003990,(Homo_sapiens:0.154176,Saimiri_sciureus:0.072750)1.000000:0.056539)1.000000:0.060601)0.953416:0.034360)1.000000:0.098444,((((Rhynchocyon_cirnei:0.122845,((Echinops_telfairi:0.117599,(Amblysomus_hottentotus:0.231515,PA44:0.000001)0.923913:0.032342)0.900362:0.032872,PA43:0.013345)0.919686:0.031630)0.996377:0.042398,((PA38:0.004263,(((Manis_pentadactyla:0.184275,Metacheiromys_marshi:0.117192)0.996377:0.084246,((PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.169768,(Hapalops_elongatus:0.162070,Tamandua_tetradactyla:0.137381)0.996377:0.075768)0.836957:0.042032)0.923188:0.045350,Orycteropus_afer:0.220292)0.824879:0.029253)0.821687:0.016334,(((PA42:0.000001,Procavia_capensis:0.000001)1.000000:0.098213,(Loxodonta_africana:0.232250,(Trichechus_manatus:0.207714,PA41:0.000001)0.260870:0.001159)0.289855:0.045642)0.817482:0.133012,PA40:0.002109)0.789130:0.021154)0.721711:0.027786)0.876107:0.065040,(((((Thomashuxleya_externa_comb:0.127444,Carodnia_vieirai:0.108771)0.500000:0.036299,Moeritherium_lyonsi:0.206572)0.514493:0.057975,((Hyopsodus_paulus:0.102966,Phenacodus_intermedius:0.087784)0.452899:0.026999,(Apheliscus_insidiosus:0.118428,(Protolipterna_ellipsodontoides:0.049059,Didolodus_multicuspis:0.076745)0.639493:0.027521)0.518116:0.018872)0.770833:0.029710)0.691770:0.036770,((Protungulatum_donnae:0.127717,Leptictis_dakotensis:0.159458)0.119565:0.026168,(Sinopa_rapax:0.095635,((Vulpavus_ovatus:0.018580,Vulpavus_profectus:0.025011)1.000000:0.064319,(Canis_lupus:0.052167,(PA21:0.000001,Felis_silvestris:0.033050)1.000000:0.059180)0.999094:0.068889)0.852355:0.023900)1.000000:0.071535)0.748965:0.027674)0.567150:0.034984,((Tribosphenomys_minutus:0.110691,(Rhombomylus_turpanensis:0.081121,(Gomphos_elkema:0.078261,(Dawsonolagus_antiquus:0.025357,Oryctolagus_cuniculus:0.099436)0.958333:0.034592)1.000000:0.065339)0.886473:0.022516)0.944293:0.032247,((Cocomys_lingchaensis:0.074266,Paramys_delicatus:0.051439)0.746377:0.025948,((Ictidomys_tridecemlineatus:0.065953,(PA28:0.000001,(Castor_canadensis:0.093053,Cavia_porcellus:0.086496)0.637681:0.013102)0.951993:0.039360)0.934783:0.044737,Rattus_norvegicus:0.139132)0.955163:0.035999)0.908213:0.026291)1.000000:0.097708)0.527576:0.032620)0.626930:0.036437,PA5:0.001318)0.627313:0.011388)0.510239:0.014105)0.934216:0.045230,PA3:0.003055)0.931522:0.029833)0.872153:0.053954)0.937198:0.075931)0.826087:0.080480)1.000000:0.100698)1.000000:0.077088,Morganucodon_watsoni:0.007896,Morganucodon_oehleri:0.008586);

["typical preservation" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: nodal values = standard bootstrap values]
(Morganucodon_watsoni:0.007851,(Haldanodon_exspectatus:0.097254,((Zhangheotherium_quinquecuspidens:0.054459,Henkelotherium_guimarotae:0.078163)0.938889:0.055612,((Tachyglossus_aculeatus:0.088633,Ornithorhynchus_anatinus:0.073670)1.000000:0.103283,(PA1:0.000001,((((Dromiciops_gliroides:0.000001,PA4:0.000001)1.000000:0.073426,(Didelphis_virginiana:0.048969,Pucadelphys_andinus:0.120006)0.994444:0.060813)1.000000:0.051689,((Eomaia_scansoria:0.065837,Maelestes_gobiensis:0.095407)0.563889:0.035150,(Zalambdalestes_lechei:0.123139,Ukhaatherium_nessovi:0.107325)0.861111:0.059198)0.777778:0.081144)0.261111:0.040535,((((((((Thomashuxleya_externa_comb:0.166412,((Mesonyx_obtusidens:0.164417,(((Artiocetus_clavis:0.118325,((Tursiops_truncatus:0.170707,Caperea_marginata:0.163982)0.997222:0.110418,Basilosaurus_cetoides:0.149532)0.569444:0.054914)0.938889:0.062215,Rodhocetus_balochistanensis:0.097270)0.494444:0.041739,(Archaeotherium_mortoni:0.138325,(Hippopotamus_amphibius:0.191090,(Sus_scrofa:0.173917,((Lama_glama:0.121758,Bos_taurus:0.124728)1.000000:0.050259,(Equus_caballus:0.131717,Mesohippus_bairdi:0.162697)0.955556:0.066380)0.408333:0.034543)0.402778:0.036463)0.355556:0.027620)0.361111:0.028614)0.197222:0.020585)0.836111:0.032859,(Carodnia_vieirai:0.086587,(Moeritherium_lyonsi:0.143174,(Trichechus_manatus:0.190621,Loxodonta_africana:0.177314)0.441667:0.034134)0.713889:0.041879)0.891667:0.069028)0.288889:0.028209)0.575000:0.031356,(((Apheliscus_insidiosus:0.112145,Protolipterna_ellipsodontoides:0.051654)0.252778:0.024925,Didolodus_multicuspis:0.078048)0.236111:0.016612,(Hyopsodus_paulus:0.102336,Phenacodus_intermedius:0.089031)0.530556:0.029482)0.594444:0.035459)0.230556:0.035168,(((Sinopa_rapax:0.088231,((Vulpavus_profectus:0.026098,Vulpavus_ovatus:0.017852)1.000000:0.061891,(Felis_silvestris:0.098672,Canis_lupus:0.065495)0.988889:0.053909)0.727778:0.027342)1.000000:0.068250,Leptictis_dakotensis:0.181046)0.141667:0.016931,Protungulatum_donnae:0.147494)0.136111:0.029057)0.088889:0.027904,(((Cocomys_lingchaensis:0.075119,Paramys_delicatus:0.051556)0.377778:0.020390,((Ictidomys_tridecemlineatus:0.064445,(Cavia_porcellus:0.090868,Castor_canadensis:0.085650)0.933333:0.049058)0.538889:0.046000,Rattus_norvegicus:0.151029)0.361111:0.027036)0.738889:0.030707,(Tribosphenomys_minutus:0.108065,(Rhombomylus_turpanensis:0.076384,(Gomphos_elkema:0.076293,(Dawsonolagus_antiquus:0.025388,Oryctolagus_cuniculus:0.100463)0.977778:0.035650)1.000000:0.070744)0.586111:0.024417)0.805556:0.034209)1.000000:0.094628)0.152778:0.032518,(Procavia_capensis:0.223448,(Rhynchocyon_cirnei:0.149062,(PA40:0.017060,(Orycteropus_afer:0.226290,(((Metacheiromys_marshi:0.118270,Manis_pentadactyla:0.186741)0.988889:0.095035,((Tamandua_tetradactyla:0.136681,Hapalops_elongatus:0.166322)1.000000:0.076927,(PA39:0.000001,Dasypus_novemcinctus:0.000001)1.000000:0.169230)0.688889:0.036372)0.850000:0.059524,PA38:0.000510)0.308333:0.013254)0.455556:0.014948)0.444444:0.027339)0.380556:0.024850)0.452778:0.056355)0.313889:0.037618,PA5:0.003486)0.233333:0.018413,(((Solenodon_paradoxus:0.174364,((Erinaceus_europaeus:0.118259,Echinops_telfairi:0.159087)0.800000:0.057435,(Sorex_araneus:0.146063,(Talpa_europaea:0.129064,Amblysomus_hottentotus:0.238535)0.477778:0.046513)0.811111:0.057858)0.608333:0.030293)0.355556:0.028476,(PA7:0.014441,(Pteropus_giganteus:0.180389,((Onychonycteris_finneyi:0.031032,Icaronycteris_index:0.026242)1.000000:0.082464,((Pteronotus_parnellii:0.075480,Myotis_lucifugus:0.062190)0.788889:0.026027,((Saccopteryx_bilineata:0.052457,Rhinopoma_hardwickii:0.051356)0.758333:0.024074,Nycteris_thebaica:0.094302)1.000000:0.032113)0.988889:0.038363)0.963889:0.041449)1.000000:0.128657)0.808333:0.031634)0.533333:0.043679,(((Cynocephalus_volans:0.015365,Galeopterus_variegatus:0.020395)1.000000:0.173141,((Ptilocercus_lowii:0.076003,Tupaia_glis:0.062794)1.000000:0.055010,(((Notharctus_tenebrosus:0.127725,Lemur_catta:0.064846)0.800000:0.036622,(Tarsius_syrichta:0.143528,(Homo_sapiens:0.156559,Saimiri_sciureus:0.072450)1.000000:0.050932)0.711111:0.025680)1.000000:0.068383,PA26:0.002558)0.455556:0.024981)0.580556:0.029334)1.000000:0.071187,PA6:0.001900)0.397222:0.026239)0.125000:0.027279)0.625000:0.043670,PA3:0.003553)0.705556:0.029699)0.438889:0.053533)0.800000:0.076437)0.636111:0.080689)1.000000:0.100607)1.000000:0.077604,Morganucodon_oehleri:0.008651);

["typical preservation" PAs corrected for the O'Leary et al. character ontology, but only PAs for nodes at the superordinal level and above included: nodal values = "transfer bootstrap expectation" values]
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