This research was supported by funding from the National Natural Science Foundation of China (U1901201), the National Key Research and Development Program of China (2020YFE0202300), the 111 Project Crop Genomics and Molecular Breeding (B20051) to J.Y., the US National Science Foundation (1546719) to J.R-I. and D.E.R, the US Dept. of Agriculture (Hatch project CA-D-PLS-2066-H 548) to J.R.-I., and the National Natural Science Foundation of China (32222062) to N.Y.