The evolved human social structure (left) of reciprocal exogamy including the exchange of mates, goods, and services (double-headed arrows), involves multiple kin lineages (filled circles) often existing in multiple residential communities (open circles).

Extensive cooperation (overlap of filled circles) likely results in economies of scale within and across human communities. In contrast, in other primates (right) one or the other sex emigrates (dotted arrows). The lack of any reciprocal exogamy means that kin lineages are isolated to single communities and thus do not generate a meta-group social structure as found in humans [2], [3].