Hierarchical levels in the additive partitioning of diversity used in the study: the diversity of each scale was linked additively to form the diversity of the next higher scale (adapted from Wagner et al., 2000; Gering et al., 2003; Chávez and Macdonald, 2012).

Hierarchical levels in the additive partitioning of diversity used in the study: the diversity of each scale was linked additively to form the diversity of the next higher scale (adapted from Wagner et al., 2000; Gering et al., 2003; Chávez and Macdonald, 2012).