Movie S4. Four types survive the initial bottleneck: CNp (orange), cNp (green), cnP (purple), and cnp (blue). CNp and cnP complement each other, forming a mutualistic coalition at low densities, but at high densities, competition from the other types causes collapse of the CNp population, nearly leading to extinction. Once densities are low enough, CNp rebounds quickly, leading to cycling. 

• cooperation
• enzymes
• microbial ecology
• public goods
• game theory

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